Author information:
(1)State Key Laboratory of Genetic Engineering and Ministry of Education Key 
Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan 
University, Shanghai, China CAS-MPG Partner Institute for Computational Biology, 
Shanghai Institutes for Biological Sciences (SIBS), Chinese Academy of Sciences 
(CAS), Shanghai, China.
(2)State Key Laboratory of Genetic Engineering and Ministry of Education Key 
Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan 
University, Shanghai, China CAS-MPG Partner Institute for Computational Biology, 
Shanghai Institutes for Biological Sciences (SIBS), Chinese Academy of Sciences 
(CAS), Shanghai, China Department of Molecular Biology and Genetics, Cornell 
University.
(3)CAS-MPG Partner Institute for Computational Biology, Shanghai Institutes for 
Biological Sciences (SIBS), Chinese Academy of Sciences (CAS), Shanghai, China.
(4)State Key Laboratory of Genetic Engineering and Ministry of Education Key 
Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan 
University, Shanghai, China CAS-MPG Partner Institute for Computational Biology, 
Shanghai Institutes for Biological Sciences (SIBS), Chinese Academy of Sciences 
(CAS), Shanghai, China lijin@fudan.edu.cn.