Author information:
(1)MOE Key Laboratory of Bioinformatics, Beijing Advanced Innovation Center for 
Structural Biology, Center for Synthetic and Systems Biology, Tsinghua-Peking 
Center for Life Sciences, School of Life Sciences, Tsinghua University, 100084, 
Beijing, China.
(2)Beijing Advanced Innovation Center for Genomics (ICG), Biomedical Pioneering 
Innovation Center (BIOPIC), Peking University, 100871, Beijing, China.
(3)State Key Laboratory of Protein and Plant Gene Research, School of Life 
Sciences, Center for Bioinformatics, Peking University, 100871, Beijing, China.
(4)Bioinformatics Division, BNRist, Department of Automation, Tsinghua 
University, 100084, Beijing, China.
(5)Department of Biological Sciences, Center for Systems Biology, The University 
of Texas, Dallas 800 West Campbell Road, RL11, Richardson, TX, 75080-3021, USA.
(6)MOE Key Laboratory of Bioinformatics, Center for Synthetic and Systems 
Biology, School of Medicine, Tsinghua University, 100084, Beijing, China.
(7)Department of Computer Science and Engineering, University of California, 
Riverside, CA, 92521, USA.
(8)Bioinformatics Division, BNRIST; Department of Computer Science and 
Technology, Tsinghua University, 100084, Beijing, China.
(9)MOE Key Laboratory of Bioinformatics, Beijing Advanced Innovation Center for 
Structural Biology, Center for Synthetic and Systems Biology, Tsinghua-Peking 
Center for Life Sciences, School of Life Sciences, Tsinghua University, 100084, 
Beijing, China. qczhang@tsinghua.edu.cn.