Author information:
(1)State Key Laboratory of Crop Biology, Shandong Agricultural University, 
Tai-An, Shandong, China; MOA Key Laboratory of Horticultural Crop Biology and 
Germplasm Innovation, Shandong Agricultural University, Tai-An, Shandong, China; 
College of Horticulture Science and Engineering, Shandong Agricultural 
University, Tai-An, Shandong, China.
(2)State Key Laboratory of Crop Biology, Shandong Agricultural University, 
Tai-An, Shandong, China; MOA Key Laboratory of Horticultural Crop Biology and 
Germplasm Innovation, Shandong Agricultural University, Tai-An, Shandong, China; 
College of Horticulture Science and Engineering, Shandong Agricultural 
University, Tai-An, Shandong, China. Electronic address: haoyujin@sdau.edu.com.
(3)State Key Laboratory of Crop Biology, Shandong Agricultural University, 
Tai-An, Shandong, China; MOA Key Laboratory of Horticultural Crop Biology and 
Germplasm Innovation, Shandong Agricultural University, Tai-An, Shandong, China; 
College of Horticulture Science and Engineering, Shandong Agricultural 
University, Tai-An, Shandong, China. Electronic address: youchunxiang@126.com.