Author information:
(1)Manchester Centre for Genomic Medicine, Evolution and Genomic Sciences, 
University of Manchester, Manchester, UK.
(2)Manchester Centre for Genomic Medicine, Evolution and Genomic Sciences, 
University of Manchester, Manchester, UK; Manchester Centre for Genomic 
Medicine, St. Mary's Hospital, Manchester University NHS Foundation Trust, 
Manchester, UK.
(3)Department of Ophthalmology, Inverclyde Royal Hospital, Greenock, PA16 0XN, 
UK.
(4)Department of Otolaryngology, Inverclyde Royal Hospital, Greenock, PA16 0XN, 
UK.
(5)Manchester Centre for Genomic Medicine, Evolution and Genomic Sciences, 
University of Manchester, Manchester, UK; Manchester Centre for Genomic 
Medicine, St. Mary's Hospital, Manchester University NHS Foundation Trust, 
Manchester, UK. Electronic address: William.newman@manchester.ac.uk.