Author information:
(1)Bioinformatics and Genomics Program, The Pennsylvania State University, 
University Park, State College, PA, 16802, USA.
(2)Department of Biochemistry and Molecular Biology, College of Medicine, The 
Pennsylvania State Hershey, Hershey, PA, 17033, USA.
(3)Department of Computer and Information Science, University of Pennsylvania, 
Philadelphia, PA, 19104, USA.
(4)Department of Genetics, The Edison Family Center for Genome Sciences and 
Systems Biology, Washington University School of Medicine, St. Louis, MO, 63108, 
USA.
(5)Department of Genetics, University of North Carolina, Chapel Hill, NC, 27599, 
USA.
(6)Department of Biostatistics, University of North Carolina, Chapel Hill, NC, 
27599, USA.
(7)Department of Computer Science, University of North Carolina, Chapel Hill, 
NC, 27599, USA.
(8)Department of Quantitative Health Sciences, Lerner Research Institute, 
Cleveland Clinic Foundation, Cleveland, OH, 44195, USA.
(9)Center for Computational Biology and Bioinformatics, Huck Institutes of the 
Life Sciences, The Pennsylvania State University, University Park, State 
College, PA, 16802, USA.
(10)Department of Genetics, The Edison Family Center for Genome Sciences and 
Systems Biology, Washington University School of Medicine, St. Louis, MO, 63108, 
USA. twang@genetics.wustl.edu.
(11)Bioinformatics and Genomics Program, The Pennsylvania State University, 
University Park, State College, PA, 16802, USA. fyue@hmc.psu.edu.
(12)Department of Biochemistry and Molecular Biology, College of Medicine, The 
Pennsylvania State Hershey, Hershey, PA, 17033, USA. fyue@hmc.psu.edu.