Author information:
(1)Center of Frontier Research, National Institute of Genetics, Research 
Organization of Information and Systems, Yata 1111, Mishima, Shizuoka 411-8540, 
Japan.
(2)Division of Biological Science, Graduate School of Science, Nagoya 
University, Chikusa-ku, Nagoya 464-8602, Japan; PRESTO, Japan Science and 
Technology Agency, 4-1-8 Honcho, Kawaguchi, Saitama 332-0012, Japan.
(3)Division of Mammalian Development, Genetic Strains Research Center, National 
Institute of Genetics, Yata 1111, Mishima, Shizuoka 411-8540, Japan; Department 
of Genetics, SOKENDAI, Yata 1111, Mishima, Shizuoka 411-8540, Japan; Department 
of Biological Sciences, Graduate School of Science, The University of Tokyo, 
7-3-1 Hongo, Bunkyo-ku, Tokyo 113-0033, Japan.
(4)Center of Frontier Research, National Institute of Genetics, Research 
Organization of Information and Systems, Yata 1111, Mishima, Shizuoka 411-8540, 
Japan; PRESTO, Japan Science and Technology Agency, 4-1-8 Honcho, Kawaguchi, 
Saitama 332-0012, Japan; Department of Genetics, SOKENDAI, Yata 1111, Mishima, 
Shizuoka 411-8540, Japan. Electronic address: mkanemak@nig.ac.jp.