document_html:
<!DOCTYPE html>
<HTML class="client-js ve-not-available" lang="en" dir="ltr"><HEAD>

<TITLE>Colony (biology) - Wikipedia</TITLE>

<LINK rel="stylesheet" href="/w/load.php?debug=false&lang=en&modules=ext.cite.styles%7Cext.uls.interlanguage%7Cext.visualEditor.desktopArticleTarget.noscript%7Cext.wikimediaBadges%7Cmediawiki.legacy.commonPrint%2Cshared%7Cmediawiki.sectionAnchor%7Cmediawiki.skinning.interface%7Cskins.vector.styles%7Cwikibase.client.init&only=styles&skin=vector" />

<STYLE>
.referencetooltip{position:absolute;list-style:none;list-style-image:none;opacity:0;font-size:10px;margin:0;z-index:5;padding:0}.referencetooltip li{border:#080086 2px solid;max-width:260px;padding:10px 8px 13px 8px;margin:0px;background-color:#F7F7F7;-webkit-box-shadow:2px 4px 2px rgba(0,0,0,0.3);-moz-box-shadow:2px 4px 2px rgba(0,0,0,0.3);box-shadow:2px 4px 2px rgba(0,0,0,0.3)}.referencetooltip li+li{margin-left:7px;margin-top:-2px;border:0;padding:0;height:3px;width:0px;background-color:transparent;-webkit-box-shadow:none;-moz-box-shadow:none;box-shadow:none;border-top:12px #080086 solid;border-right:7px transparent solid;border-left:7px transparent solid}.referencetooltip>li+li::after{content:'';border-top:8px #F7F7F7 solid;border-right:5px transparent solid;border-left:5px transparent solid;margin-top:-12px;margin-left:-5px;z-index:1;height:0px;width:0px;display:block}.client-js body .referencetooltip li li{border:none;-webkit-box-shadow:none;-moz-box-shadow:none;box-shadow:none;height:auto;width:auto;margin:auto;padding:0;position:static}.RTflipped{padding-top:13px}.referencetooltip.RTflipped li+li{position:absolute;top:2px;border-top:0;border-bottom:12px #080086 solid}.referencetooltip.RTflipped li+li::after{border-top:0;border-bottom:8px #F7F7F7 solid;position:absolute;margin-top:7px}.RTsettings{float:right;height:24px;width:24px;cursor:pointer;background-image:url(//upload.wikimedia.org/wikipedia/commons/thumb/0/05/OOjs_UI_icon_advanced.svg/24px-OOjs_UI_icon_advanced.svg.png);background-image:linear-gradient(transparent,transparent),url(//upload.wikimedia.org/wikipedia/commons/0/05/OOjs_UI_icon_advanced.svg);margin-top:-9px;margin-right:-7px;-webkit-transition:opacity 0.15s;-moz-transition:opacity 0.15s;-ms-transition:opacity 0.15s;-o-transition:opacity 0.15s;transition:opacity 0.15s;opacity:0.6;filter:alpha(opacity=60)}.RTsettings:hover{opacity:1;filter:alpha(opacity=100)}.RTTarget{border:#080086 2px solid}
.wp-teahouse-question-form{position:absolute;margin-left:auto;margin-right:auto;background-color:#f4f3f0;border:1px solid #a7d7f9;padding:1em}#wp-th-question-ask{float:right}.wp-teahouse-ask a.external{background-image:none !important}.wp-teahouse-respond-form{position:absolute;margin-left:auto;margin-right:auto;background-color:#f4f3f0;border:1px solid #a7d7f9;padding:1em}.wp-th-respond{float:right}.wp-teahouse-respond a.external{background-image:none !important}
.cite-accessibility-label{ top:-99999px;clip:rect(1px 1px 1px 1px); clip:rect(1px,1px,1px,1px); position:absolute !important;padding:0 !important;border:0 !important;height:1px !important;width:1px !important; overflow:hidden}
.mw-spinner{background-color:transparent;background-position:center center;background-repeat:no-repeat}.mw-spinner-small{background-image:url(data:image/gif;base64,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);background-image:url(/w/resources/src/jquery/images/spinner.gif?ca65b)!ie;height:20px;width:20px; min-width:20px}.mw-spinner-large{background-image:url(data:image/gif;base64,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);background-image:url(/w/resources/src/jquery/images/spinner-large.gif?57f34)!ie;height:32px;width:32px; min-width:32px}.mw-spinner-block{display:block; width:100%}.mw-spinner-inline{display:inline-block;vertical-align:middle; zoom:1;*display:inline; }
.suggestions{overflow:hidden;position:absolute;top:0;left:0;width:0;border:0;z-index:1099;padding:0;margin:-1px 0 0 0}.suggestions-special{position:relative;background-color:#fff;cursor:pointer;border:1px solid #a2a9b1;margin:0;margin-top:-2px;display:none;padding:0.25em 0.25em;line-height:1.25em}.suggestions-results{background-color:#fff;cursor:pointer;border:1px solid #a2a9b1;padding:0;margin:0}.suggestions-result{color:#000;margin:0;line-height:1.5em;padding:0.01em 0.25em;text-align:left; overflow:hidden;text-overflow:ellipsis;white-space:nowrap}.suggestions-result-current{background-color:#2a4b8d;color:#fff}.suggestions-special .special-label{color:#72777d;text-align:left}.suggestions-special .special-query{color:#000;font-style:italic;text-align:left}.suggestions-special .special-hover{background-color:#c8ccd1}.suggestions-result-current .special-label,.suggestions-result-current .special-query{color:#fff}.highlight{font-weight:bold}
@media screen {
	.tochidden,.toctoggle{-moz-user-select:none;-webkit-user-select:none;-ms-user-select:none;user-select:none}.toctoggle{font-size:94%}}
@media print {
	.toc.tochidden,.toctoggle{display:none}}
.uls-menu{border-radius:2px; font-size:medium}.uls-search,.uls-language-settings-close-block{border-top-right-radius:2px;border-top-left-radius:2px}.uls-language-list{border-bottom-right-radius:2px;border-bottom-left-radius:2px}.uls-menu.callout:before,.uls-menu.callout:after{border-top:10px solid transparent;border-bottom:10px solid transparent;display:inline-block; top:17px;position:absolute;content:''}.uls-menu.callout.selector-right:before{ border-left:10px solid #c8ccd1; right:-11px}.uls-menu.callout.selector-right:after{ border-left:10px solid #f8f9fa; right:-10px}.uls-menu.callout.selector-left:before{ border-right:10px solid #c8ccd1; left:-11px}.uls-menu.callout.selector-left:after{ border-right:10px solid #f8f9fa; left:-10px}.uls-ui-languages button{margin:5px 15px 5px 0;white-space:nowrap;overflow:hidden}.uls-search-wrapper-wrapper{position:relative;padding-left:40px;margin-top:5px;margin-bottom:5px}.uls-icon-back{background:transparent url(/w/extensions/UniversalLanguageSelector/resources/images/back-grey-ltr.png?90e9b) no-repeat scroll center center;background-image:-webkit-linear-gradient(transparent,transparent),url(/w/extensions/UniversalLanguageSelector/resources/images/back-grey-ltr.svg?e226b);background-image:linear-gradient(transparent,transparent),url("data:image/svg+xml,%3Csvg xmlns=%22http://www.w3.org/2000/svg%22 width=%2224%22 height=%2224%22 viewBox=%220 0 24 24%22%3E %3Cpath fill=%22%2354595d%22 d=%22M7 13.1l8.9 8.9c.8-.8.8-2 0-2.8l-6.1-6.1 6-6.1c.8-.8.8-2 0-2.8L7 13.1z%22/%3E %3C/svg%3E");background-size:28px;background-position:center center;height:32px;width:40px;display:block;position:absolute;left:0;border-right:1px solid #c8ccd1;opacity:0.8}.uls-icon-back:hover{opacity:1;cursor:pointer}.uls-menu .uls-no-results-view .uls-no-found-more{background-color:#fff}.uls-menu .uls-no-results-view h3{padding:0 28px;margin:0;color:#54595d;font-size:1em;font-weight:normal}  .skin-vector .uls-menu{border-color:#c8ccd1;-webkit-box-shadow:0 2px 2px 0 rgba(0,0,0,0.25);box-shadow:0 2px 2px 0 rgba(0,0,0,0.25);font-size:0.875em}.skin-vector .uls-search{border-bottom-color:#c8ccd1}.skin-vector .uls-filtersuggestion{color:#72777d}.skin-vector .uls-lcd-region-title{color:#54595d}
.mw-editfont-monospace{font-family:monospace,monospace}.mw-editfont-sans-serif{font-family:sans-serif}.mw-editfont-serif{font-family:serif} .mw-editfont-monospace,.mw-editfont-sans-serif,.mw-editfont-serif{font-size:13px}
@-webkit-keyframes centralAuthPPersonalAnimation{0%{opacity:0;-webkit-transform:translateY(-20px)}100%{opacity:1;-webkit-transform:translateY(0)}}@-moz-keyframes centralAuthPPersonalAnimation{0%{opacity:0;-moz-transform:translateY(-20px)}100%{opacity:1;-moz-transform:translateY(0)}}@-o-keyframes centralAuthPPersonalAnimation{0%{opacity:0;-o-transform:translateY(-20px)}100%{opacity:1;-o-transform:translateY(0)}}@keyframes centralAuthPPersonalAnimation{0%{opacity:0;transform:translateY(-20px)}100%{opacity:1;transform:translateY(0)}}.centralAuthPPersonalAnimation{-webkit-animation-duration:1s;-moz-animation-duration:1s;-o-animation-duration:1s;animation-duration:1s;-webkit-animation-fill-mode:both;-moz-animation-fill-mode:both;-o-animation-fill-mode:both;animation-fill-mode:both;-webkit-animation-name:centralAuthPPersonalAnimation;-moz-animation-name:centralAuthPPersonalAnimation;-o-animation-name:centralAuthPPersonalAnimation;animation-name:centralAuthPPersonalAnimation}
@media print{#centralNotice{display:none}}.cn-closeButton{display:inline-block;zoom:1;background:url(data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAABQAAAAUBAMAAAB/pwA+AAAAElBMVEUAAAAQEBDPz88AAABAQEDv7+9oe1vvAAAABnRSTlMA3rLe3rJS22KzAAAARElEQVQI12PAAUIUQCSTK5BwFgIxFU1AhKECUFAYKAAioXwwBeZChMGCEGGQIFQYJohgIhQgtCEMQ7ECYTHCOciOxA4AADgJTXIb9s8AAAAASUVORK5CYII=) no-repeat;background:url(/w/extensions/CentralNotice/resources/subscribing/close.png?8e3d8) no-repeat!ie;width:20px;height:20px;text-indent:20px;white-space:nowrap;overflow:hidden}
.mw-ui-button{font-family:inherit;font-size:1em;display:inline-block;min-width:4em;max-width:28.75em;padding:0.546875em 1em;line-height:1.286;margin:0;border-radius:2px;-webkit-box-sizing:border-box;-moz-box-sizing:border-box;box-sizing:border-box;-webkit-appearance:none;*display:inline; zoom:1;vertical-align:middle;background-color:#f8f9fa;color:#222222;border:1px solid #a2a9b1;text-align:center;font-weight:bold;cursor:pointer}.mw-ui-button:visited{color:#222222}.mw-ui-button:hover{background-color:#ffffff;color:#444444;border-color:#a2a9b1}.mw-ui-button:focus{background-color:#ffffff;color:#222222;border-color:#3366cc;box-shadow:inset 0 0 0 1px #3366cc,inset 0 0 0 2px #ffffff}.mw-ui-button:active,.mw-ui-button.is-on,.mw-ui-button.mw-ui-checked{background-color:#d9d9d9;color:#000000;border-color:#72777d;box-shadow:none}.mw-ui-button:disabled{background-color:#c8ccd1;color:#fff;border-color:#c8ccd1}.mw-ui-button:disabled:hover,.mw-ui-button:disabled:active{background-color:#c8ccd1;color:#fff;box-shadow:none;border-color:#c8ccd1}.mw-ui-button:focus{outline-width:0}.mw-ui-button:focus::-moz-focus-inner{border-color:transparent;padding:0}.mw-ui-button:not(:disabled){-webkit-transition:background-color 100ms,color 100ms,border-color 100ms,box-shadow 100ms;-moz-transition:background-color 100ms,color 100ms,border-color 100ms,box-shadow 100ms;transition:background-color 100ms,color 100ms,border-color 100ms,box-shadow 100ms}.mw-ui-button:disabled{text-shadow:none;cursor:default}.mw-ui-button.mw-ui-big{font-size:1.3em}.mw-ui-button.mw-ui-block{display:block;width:100%;margin-left:auto;margin-right:auto}.mw-ui-button.mw-ui-progressive{background-color:#3366cc;color:#fff;border:1px solid #3366cc}.mw-ui-button.mw-ui-progressive:hover{background-color:#447ff5;border-color:#447ff5}.mw-ui-button.mw-ui-progressive:focus{box-shadow:inset 0 0 0 1px #3366cc,inset 0 0 0 2px #ffffff}.mw-ui-button.mw-ui-progressive:active,.mw-ui-button.mw-ui-progressive.is-on,.mw-ui-button.mw-ui-progressive.mw-ui-checked{background-color:#2a4b8d;border-color:#2a4b8d;box-shadow:none}.mw-ui-button.mw-ui-progressive:disabled{background-color:#c8ccd1;color:#fff;border-color:#c8ccd1}.mw-ui-button.mw-ui-progressive:disabled:hover,.mw-ui-button.mw-ui-progressive:disabled:active,.mw-ui-button.mw-ui-progressive:disabled.mw-ui-checked{background-color:#c8ccd1;color:#fff;border-color:#c8ccd1;box-shadow:none}.mw-ui-button.mw-ui-progressive.mw-ui-quiet{color:#222222}.mw-ui-button.mw-ui-progressive.mw-ui-quiet:hover{background-color:transparent;color:#447ff5}.mw-ui-button.mw-ui-progressive.mw-ui-quiet:active,.mw-ui-button.mw-ui-progressive.mw-ui-quiet.mw-ui-checked{color:#2a4b8d}.mw-ui-button.mw-ui-progressive.mw-ui-quiet:focus{background-color:transparent;color:#3366cc}.mw-ui-button.mw-ui-progressive.mw-ui-quiet:disabled{color:#c8ccd1}.mw-ui-button.mw-ui-destructive{background-color:#dd3333;color:#fff;border:1px solid #dd3333}.mw-ui-button.mw-ui-destructive:hover{background-color:#ff4242;border-color:#ff4242}.mw-ui-button.mw-ui-destructive:focus{box-shadow:inset 0 0 0 1px #dd3333,inset 0 0 0 2px #ffffff}.mw-ui-button.mw-ui-destructive:active,.mw-ui-button.mw-ui-destructive.is-on,.mw-ui-button.mw-ui-destructive.mw-ui-checked{background-color:#b32424;border-color:#b32424;box-shadow:none}.mw-ui-button.mw-ui-destructive:disabled{background-color:#c8ccd1;color:#fff;border-color:#c8ccd1}.mw-ui-button.mw-ui-destructive:disabled:hover,.mw-ui-button.mw-ui-destructive:disabled:active,.mw-ui-button.mw-ui-destructive:disabled.mw-ui-checked{background-color:#c8ccd1;color:#fff;border-color:#c8ccd1;box-shadow:none}.mw-ui-button.mw-ui-destructive.mw-ui-quiet{color:#222222}.mw-ui-button.mw-ui-destructive.mw-ui-quiet:hover{background-color:transparent;color:#ff4242}.mw-ui-button.mw-ui-destructive.mw-ui-quiet:active,.mw-ui-button.mw-ui-destructive.mw-ui-quiet.mw-ui-checked{color:#b32424}.mw-ui-button.mw-ui-destructive.mw-ui-quiet:focus{background-color:transparent;color:#dd3333}.mw-ui-button.mw-ui-destructive.mw-ui-quiet:disabled{color:#c8ccd1}.mw-ui-button.mw-ui-quiet{background:transparent;border:0;text-shadow:none;color:#222222}.mw-ui-button.mw-ui-quiet:hover{background-color:transparent;color:#444444}.mw-ui-button.mw-ui-quiet:active,.mw-ui-button.mw-ui-quiet.mw-ui-checked{color:#000000}.mw-ui-button.mw-ui-quiet:focus{background-color:transparent;color:#222222}.mw-ui-button.mw-ui-quiet:disabled{color:#c8ccd1}.mw-ui-button.mw-ui-quiet:hover,.mw-ui-button.mw-ui-quiet:focus{box-shadow:none}.mw-ui-button.mw-ui-quiet:active,.mw-ui-button.mw-ui-quiet:disabled{background:transparent}input.mw-ui-button::-moz-focus-inner,button.mw-ui-button::-moz-focus-inner{margin-top:-1px;margin-bottom:-1px}a.mw-ui-button{text-decoration:none}a.mw-ui-button:hover,a.mw-ui-button:focus{text-decoration:none}.mw-ui-button-group > *{min-width:48px;border-radius:0;float:left}.mw-ui-button-group > *:first-child{border-top-left-radius:2px;border-bottom-left-radius:2px}.mw-ui-button-group > *:not(:first-child){border-left:0}.mw-ui-button-group > *:last-child{border-top-right-radius:2px;border-bottom-right-radius:2px}.mw-ui-button-group .is-on .button{cursor:default}
.mw-ui-icon{position:relative;line-height:1.5em;min-height:1.5em;min-width:1.5em}.mw-ui-icon.mw-ui-icon-element{text-indent:-999px;overflow:hidden;width:3.5em;min-width:3.5em;max-width:3.5em}.mw-ui-icon.mw-ui-icon-element:before{left:0;right:0;position:absolute;margin:0 1em}.mw-ui-icon.mw-ui-icon-element.mw-ui-icon-large{width:4.625em;min-width:4.625em;max-width:4.625em;line-height:4.625em;min-height:4.625em}.mw-ui-icon.mw-ui-icon-element.mw-ui-icon-large:before{min-height:4.625em}.mw-ui-icon.mw-ui-icon-before:before,.mw-ui-icon.mw-ui-icon-element:before{background-position:50% 50%;background-repeat:no-repeat;background-size:100% auto;float:left;display:block;min-height:1.5em;content:''}.mw-ui-icon.mw-ui-icon-before:before{position:relative;width:1.5em;margin-right:1em}.mw-ui-icon.mw-ui-icon-small:before{background-size:66.67% auto}
.mw-ui-icon-popups-settings:before{background-image:url(/w/load.php?modules=ext.popups.images&image=popups-settings&format=rasterized&lang=en&skin=vector&version=0l3es32);background-image:linear-gradient(transparent,transparent),url("data:image/svg+xml,%3Csvg xmlns=%22http://www.w3.org/2000/svg%22 viewBox=%220 0 24 24%22%3E %3Cpath fill=%22%2354595d%22 d=%22M20 14.5v-2.9l-1.8-.3c-.1-.4-.3-.8-.6-1.4l1.1-1.5-2.1-2.1-1.5 1.1c-.5-.3-1-.5-1.4-.6L13.5 5h-2.9l-.3 1.8c-.5.1-.9.3-1.4.6L7.4 6.3 5.3 8.4l1 1.5c-.3.5-.4.9-.6 1.4l-1.7.2v2.9l1.8.3c.1.5.3.9.6 1.4l-1 1.5 2.1 2.1 1.5-1c.4.2.9.4 1.4.6l.3 1.8h3l.3-1.8c.5-.1.9-.3 1.4-.6l1.5 1.1 2.1-2.1-1.1-1.5c.3-.5.5-1 .6-1.4l1.5-.3zM12 16c-1.7 0-3-1.3-3-3s1.3-3 3-3 3 1.3 3 3-1.3 3-3 3z%22/%3E %3C/svg%3E")}.mw-ui-icon-popups-close:before{background-image:url(/w/load.php?modules=ext.popups.images&image=popups-close&format=rasterized&lang=en&skin=vector&version=0l3es32);background-image:linear-gradient(transparent,transparent),url("data:image/svg+xml,%3Csvg xmlns=%22http://www.w3.org/2000/svg%22 width=%2220%22 height=%2220%22 viewBox=%220 0 20 20%22%3E %3Cpath d=%22M3.636 2.222l14.142 14.142-1.414 1.414L2.222 3.636z%22/%3E %3Cpath d=%22M17.778 3.636L3.636 17.778l-1.414-1.414L16.364 2.222z%22/%3E %3C/svg%3E")}.mw-ui-icon-preview-generic:before{background-image:url(/w/load.php?modules=ext.popups.images&image=preview-generic&format=rasterized&lang=en&skin=vector&version=0l3es32);background-image:linear-gradient(transparent,transparent),url("data:image/svg+xml,%3Csvg width=%2237%22 height=%2227%22 xmlns=%22http://www.w3.org/2000/svg%22%3E %3Cg id=%22Page-1%22 fill=%22none%22 fill-rule=%22evenodd%22%3E %3Cg id=%22sad-face%22 fill=%22%23C8CCD1%22%3E %3Cpath d=%22M5.475.7v20.075L0 26.25h31.025c3.102 0 5.475-2.372 5.475-5.475V.7H5.475zm20.44 4.562c1.277 0 2.19 1.095 2.19 2.19 0 1.096-.913 2.373-2.19 2.373-1.278 0-2.19-1.095-2.19-2.19s1.095-2.373 2.19-2.373zm-9.855 0c1.277 0 2.19 1.095 2.19 2.19 0 1.096-1.095 2.373-2.19 2.373s-2.19-1.095-2.19-2.19.913-2.373 2.19-2.373zm4.928 8.213c-7.153 0-8.415 7.012-8.415 7.012s2.805-1.403 8.415-1.403c5.61 0 8.414 1.403 8.414 1.403S28 13.475 20.988 13.475z%22 id=%22Shape%22/%3E %3C/g%3E %3C/g%3E %3C/svg%3E")}.mw-ui-icon-preview-disambiguation:before{background-image:url(/w/load.php?modules=ext.popups.images&image=preview-disambiguation&format=rasterized&lang=en&skin=vector&version=0l3es32);background-image:linear-gradient(transparent,transparent),url("data:image/svg+xml,%3Csvg xmlns=%22http://www.w3.org/2000/svg%22 width=%2220%22 height=%2220%22 viewBox=%222 2 20 20%22%3E %3Cpath fill=%22%23C8CCD1%22 d=%22M11 12h4V7h-4v5zm-5 2h9v-1H6v1zm0 2h9v-1H6v1zm0 2h9v-1H6v1zm4-9H6v1h4V9zm0 2H6v1h4v-1zm0-4H6v1h4V7zM4 5h13v16H7c-1.7 0-3-1.3-3-3V5z%22/%3E %3Cpath fill-rule=%22evenodd%22 fill=%22%23C8CCD1%22 d=%22M18 4v14h2V2H7v2%22/%3E %3C/svg%3E")}</STYLE><STYLE>
.mw-3d-wrapper{display:inline-block;position:relative;overflow:hidden;vertical-align:top}.mw-3d-badge{position:absolute;top:11px;left:11px;color:#1e1f21;font-size:14px;line-height:19px;font-weight:bold;opacity:0.8;padding:2px 5px;background-color:#f8f9fa;border-radius:2px}.mw-3d-thumb-placeholder{display:inline-block;text-decoration:none;color:#222}
.suggestions a.mw-searchSuggest-link,.suggestions a.mw-searchSuggest-link:hover,.suggestions a.mw-searchSuggest-link:active,.suggestions a.mw-searchSuggest-link:focus{color:#000;text-decoration:none}.suggestions-result-current a.mw-searchSuggest-link,.suggestions-result-current a.mw-searchSuggest-link:hover,.suggestions-result-current a.mw-searchSuggest-link:active,.suggestions-result-current a.mw-searchSuggest-link:focus{color:#fff}.suggestions a.mw-searchSuggest-link .special-query{ overflow:hidden;text-overflow:ellipsis;white-space:nowrap}
#uls-settings-block{background-color:#f8f9fa;border-top:1px solid #c8ccd1;padding-left:10px;line-height:1.2em;border-radius:0 0 2px 2px}#uls-settings-block > button{background:left top transparent no-repeat;background-size:20px auto;color:#54595d;display:inline-block;margin:8px 15px;border:0;padding:0 0 0 26px;font-size:medium;cursor:pointer}#uls-settings-block > button:hover{color:#222}#uls-settings-block > button.display-settings-block{background-image:url(/w/extensions/UniversalLanguageSelector/resources/images/display.png?d25f1);background-image:linear-gradient(transparent,transparent),url("data:image/svg+xml,%3Csvg xmlns=%22http://www.w3.org/2000/svg%22 width=%2220%22 height=%2220%22 viewBox=%220 0 20 20%22%3E %3Cpath fill=%22%23222%22 d=%22M.002 2.275v12.945h8.405c.535 1.624-.975 1.786-1.902 2.505 0 0 2.293-.024 3.439-.024 1.144 0 3.432.024 3.432.024-.905-.688-2.355-.868-1.902-2.505h8.527v-12.945h-20zm6.81 1.84h.797l3.313 8.466h-1.043l-1.043-2.638h-3.374l-1.043 2.638h-.982zm.368 1.104c-.084.369-.211.785-.368 1.227l-.982 2.577h2.699l-.982-2.577c-.128-.33-.234-.747-.368-1.227zm7.117.982c.753 0 1.295.157 1.656.491.365.334.552.858.552 1.595v4.294h-.675l-.184-.859h-.062c-.315.396-.605.655-.92.798-.311.138-.758.184-1.227.184-.626 0-1.115-.168-1.472-.491-.353-.323-.491-.754-.491-1.35 0-1.275 1.028-1.963 3.068-2.025h1.043v-.429c0-.495-.091-.87-.307-1.104-.211-.238-.574-.307-1.043-.307-.526 0-1.115.107-1.779.429l-.307-.675c.311-.169.674-.333 1.043-.429.373-.096.732-.123 1.104-.123zm.307 3.313c-.761.027-1.318.157-1.656.368-.334.207-.491.54-.491.982 0 .346.1.617.307.798.211.181.544.245.92.245.595 0 1.012-.164 1.35-.491.342-.326.552-.762.552-1.35v-.552z%22/%3E %3C/svg%3E")}#uls-settings-block > button.input-settings-block{background-image:url(/w/extensions/UniversalLanguageSelector/resources/images/input.png?aea9e);background-image:linear-gradient(transparent,transparent),url("data:image/svg+xml,%3Csvg xmlns=%22http://www.w3.org/2000/svg%22 width=%2220%22 height=%2220%22 viewBox=%220 0 20 20%22%3E %3Cpath fill=%22%23222%22 d=%22M9 1.281c-.124.259-.185.599-.5.688-.55.081-1.133.018-1.688 0-.866-.032-1.733-.148-2.594 0-.588.157-.953.727-1.188 1.25-.178.416-.271.836-.344 1.281h-2.688v11.5h20v-11.5h-16.344c.109-.52.203-1.057.563-1.469.222-.231.587-.17.875-.188 1.212.003 2.415.179 3.625.063.463-.058.812-.455.969-.875l.188-.438-.875-.313zm-7.125 5.844h1.563c.094 0 .188.093.188.188v1.531c0 .094-.093.188-.188.188h-1.563c-.094 0-.156-.093-.156-.188v-1.531c0-.094.062-.188.156-.188zm2.844 0h1.563c.094 0 .156.093.156.188v1.531c0 .094-.062.188-.156.188h-1.563c-.094 0-.156-.093-.156-.188v-1.531c0-.094.062-.188.156-.188zm2.844 0h1.563c.094 0 .156.093.156.188v1.531c0 .094-.062.188-.156.188h-1.563c-.094 0-.188-.093-.188-.188v-1.531c0-.094.093-.188.188-.188zm2.813 0h1.563c.094 0 .188.093.188.188v1.531c0 .094-.093.188-.188.188h-1.563c-.094 0-.156-.093-.156-.188v-1.531c0-.094.062-.188.156-.188zm2.844 0h1.563c.094 0 .156.093.156.188v1.531c0 .094-.062.188-.156.188h-1.563c-.094 0-.156-.093-.156-.188v-1.531c0-.094.062-.188.156-.188zm2.844 0h1.531c.094 0 .188.093.188.188v1.531c0 .094-.093.188-.188.188h-1.531c-.094 0-.188-.093-.188-.188v-1.531c0-.094.093-.188.188-.188zm-12.844 3h1.563c.094 0 .156.093.156.188v1.563c0 .094-.062.156-.156.156h-1.563c-.094 0-.156-.062-.156-.156v-1.563c0-.094.062-.188.156-.188zm2.906 0h1.563c.094 0 .188.093.188.188v1.563c0 .094-.093.156-.188.156h-1.563c-.094 0-.156-.062-.156-.156v-1.563c0-.094.062-.188.156-.188zm2.938 0h1.531c.094 0 .188.093.188.188v1.563c0 .094-.093.156-.188.156h-1.531c-.094 0-.188-.062-.188-.156v-1.563c0-.094.093-.188.188-.188zm2.906 0h1.563c.094 0 .156.093.156.188v1.563c0 .094-.062.156-.156.156h-1.563c-.094 0-.188-.062-.188-.156v-1.563c0-.094.093-.188.188-.188zm2.906 0h1.563c.094 0 .156.093.156.188v1.563c0 .094-.062.156-.156.156h-1.563c-.094 0-.156-.062-.156-.156v-1.563c0-.094.062-.188.156-.188zm-10.875 3.563h12c.088 0 .156.068.156.156v.844c0 .088-.068.156-.156.156h-12c-.088 0-.156-.068-.156-.156v-.844c0-.088.068-.156.156-.156z%22/%3E %3C/svg%3E")}
.ve-init-mw-tempWikitextEditorWidget{border:0;padding:0;color:inherit;line-height:1.5em; }.ve-init-mw-tempWikitextEditorWidget:focus{outline:0;padding:0}.ve-init-mw-tempWikitextEditorWidget::selection{background:rgba(109,169,247,0.5); }
#p-lang .body ul .uls-trigger,#p-lang .pBody ul .uls-trigger{background-image:none;padding:0} .mw-interlanguage-selector,.mw-interlanguage-selector:active{cursor:pointer;padding:4px 6px 4px 25px;font-size:13px;font-weight:normal;background-image:url(/w/extensions/UniversalLanguageSelector/resources/images/compact-links-trigger.png?b0c8e);background-image:linear-gradient(transparent,transparent),url("data:image/svg+xml,%3Csvg xmlns=%22http://www.w3.org/2000/svg%22 width=%2218%22 height=%2218%22 viewBox=%220 0 24 24%22%3E %3Cpath fill=%22%2372777d%22 d=%22M13 19l.8-3h5.3l.9 3h2.2l-4.2-13h-3l-4.2 13h2.2zm3.5-11l2 6h-4l2-6zM5 4l.938 1.906h-4.938v2.094h1.594c.6 1.8 1.406 3.206 2.406 4.406-1.1.7-4.313 1.781-4.313 1.781l1.313 1.813s3.487-1.387 4.688-2.188c1 .7 2.319 1.188 3.719 1.688l.594-2c-1-.3-1.988-.688-2.688-1.188 1.1-1.1 1.9-2.506 2.5-4.406h2.188l.5-2h-5.563l-.938-1.906h-2zm-.188 4h3.781c-.4 1.3-.906 2-1.906 3-1.1-1-1.475-1.7-1.875-3z%22/%3E %3C/svg%3E");background-size:18px;background-repeat:no-repeat;background-position:left 4px center;margin:4px 0;text-align:left}.mw-interlanguage-selector:active,.mw-interlanguage-selector.selector-open{background-color:#c8ccd1;color:#54595d}.interlanguage-uls-menu:before,.interlanguage-uls-menu:after{border-top:10px solid transparent;border-bottom:10px solid transparent;display:inline-block; top:17px;position:absolute;content:''}.interlanguage-uls-menu.selector-right:before{ border-left:10px solid #c8ccd1; right:-11px}.interlanguage-uls-menu.selector-right:after{ border-left:10px solid #f8f9fa; right:-10px}.interlanguage-uls-menu.selector-left:before{ border-right:10px solid #c8ccd1; left:-11px}.interlanguage-uls-menu.selector-left:after{ border-right:10px solid #f8f9fa; left:-10px}
.mw-mmv-overlay{position:fixed;top:0;left:0;right:0;bottom:0;z-index:1000;background-color:#000}body.mw-mmv-lightbox-open{overflow-y:auto;  }body.mw-mmv-lightbox-open #mw-page-base,body.mw-mmv-lightbox-open #mw-head-base,body.mw-mmv-lightbox-open #mw-navigation,body.mw-mmv-lightbox-open #content,body.mw-mmv-lightbox-open #footer,body.mw-mmv-lightbox-open #globalWrapper{ display:none}body.mw-mmv-lightbox-open > *{ display:none}body.mw-mmv-lightbox-open > .mw-mmv-overlay,body.mw-mmv-lightbox-open > .mw-mmv-wrapper{display:block}.mw-mmv-filepage-buttons{margin-top:5px}.mw-mmv-filepage-buttons .mw-mmv-view-expanded,.mw-mmv-filepage-buttons .mw-mmv-view-config{display:block;line-height:inherit}.mw-mmv-filepage-buttons .mw-mmv-view-expanded.mw-ui-icon:before{background-image:url("data:image/svg+xml,%3Csvg xmlns=%22http://www.w3.org/2000/svg%22 viewBox=%220 0 1024 768%22%3E %3Cpath d=%22M851.2 71.6L690.7 232.1l-40.1-40.3-9.6 164.8 164.8-9.3-40.3-40.4L926 146.4l58.5 58.5L997.6 0 792.7 13.1%22/%3E %3Cpath d=%22M769.6 89.3H611.9l70.9 70.8 7.9 7.5m-47.1 234.6l-51.2 3 3-51.2 9.4-164.4 5.8-100.3H26.4V768h883.1V387l-100.9 5.8-165 9.4zM813.9 678H113.6l207.2-270.2 31.5-12.9L548 599.8l105.9-63.2 159.8 140.8.2.6zm95.6-291.9V228l-79.1 78.9 7.8 7.9%22/%3E %3C/svg%3E")}.mw-mmv-filepage-buttons .mw-mmv-view-config.mw-ui-icon:before{background-image:url("data:image/svg+xml,%3Csvg xmlns=%22http://www.w3.org/2000/svg%22 viewBox=%220 0 1024 768%22%3E %3Cpath d=%22M897 454.6V313.4L810.4 299c-6.4-23.3-16-45.7-27.3-65.8l50.5-71.4-99.4-100.2-71.4 50.5c-20.9-11.2-42.5-20.9-65.8-27.3L582.6-1H441.4L427 85.6c-23.3 6.4-45.7 16-65.8 27.3l-71.4-50.5-100.3 99.5 50.5 71.4c-11.2 20.9-20.9 42.5-27.3 66.6L127 313.4v141.2l85.8 14.4c6.4 23.3 16 45.7 27.3 66.6L189.6 607l99.5 99.5 71.4-50.5c20.9 11.2 42.5 20.9 66.6 27.3l14.4 85.8h141.2l14.4-86.6c23.3-6.4 45.7-16 65.8-27.3l71.4 50.5 99.5-99.5-50.5-71.4c11.2-20.9 20.9-42.5 27.3-66.6l86.4-13.6zm-385 77c-81.8 0-147.6-66.6-147.6-147.6 0-81.8 66.6-147.6 147.6-147.6S659.6 302.2 659.6 384 593.8 531.6 512 531.6z%22/%3E %3C/svg%3E");opacity:0.75}.mw-mmv-filepage-buttons .mw-mmv-view-config.mw-ui-icon:before:hover{opacity:1}
#mwe-popups-svg{position:absolute;top:-1000px}.mwe-popups-settings-icon{border-radius:2px;font-size:16px; margin-right:-0.75em}.mwe-popups-settings-icon:hover{background-color:#eaecf0}.mwe-popups-settings-icon:active{background-color:#c8ccd1}.mwe-popups{background:#fff;position:absolute;z-index:110;-webkit-box-shadow:0 30px 90px -20px rgba(0,0,0,0.3),0 0 1px #a2a9b1;box-shadow:0 30px 90px -20px rgba(0,0,0,0.3),0 0 1px #a2a9b1;padding:0;display:none;font-size:14px;line-height:20px;min-width:300px;border-radius:2px; }.mwe-popups .mw-ui-icon{font-size:16px;margin:21px 0 8px 0}.mwe-popups .mwe-popups-container{color:#222222;margin-top:-9px;padding-top:9px;text-decoration:none}.mwe-popups .mwe-popups-container footer{padding:16px;margin:0;font-size:10px;position:absolute;bottom:0; left:0}.mwe-popups .mwe-popups-container footer .mwe-popups-settings-icon{float:right}.mwe-popups .mwe-popups-title{display:block;font-weight:bold;margin:0 16px}.mwe-popups .mwe-popups-extract{margin:16px;display:block;color:#222222;text-decoration:none;position:relative;   }.mwe-popups .mwe-popups-extract:hover{text-decoration:none}.mwe-popups .mwe-popups-extract:after{content:' ';position:absolute;bottom:0;width:25%;height:20px;background-color:transparent}.mwe-popups .mwe-popups-extract[dir='ltr']:after{ right:0; background-image:-webkit-linear-gradient(to right,rgba(255,255,255,0),#ffffff 50%); background-image:-moz-linear-gradient(to right,rgba(255,255,255,0),#ffffff 50%); background-image:-o-linear-gradient(to right,rgba(255,255,255,0),#ffffff 50%); background-image:linear-gradient(to right,rgba(255,255,255,0),#ffffff 50%)}.mwe-popups .mwe-popups-extract[dir='rtl']:after{ left:0; background-image:-webkit-linear-gradient(to left,rgba(255,255,255,0),#ffffff 50%); background-image:-moz-linear-gradient(to left,rgba(255,255,255,0),#ffffff 50%); background-image:-o-linear-gradient(to left,rgba(255,255,255,0),#ffffff 50%); background-image:linear-gradient(to left,rgba(255,255,255,0),#ffffff 50%)}.mwe-popups .mwe-popups-extract p{margin:0}.mwe-popups .mwe-popups-extract ul,.mwe-popups .mwe-popups-extract ol,.mwe-popups .mwe-popups-extract li,.mwe-popups .mwe-popups-extract dl,.mwe-popups .mwe-popups-extract dd,.mwe-popups .mwe-popups-extract dt{margin-top:0;margin-bottom:0}.mwe-popups svg{overflow:hidden}.mwe-popups.mwe-popups-is-tall{width:450px}.mwe-popups.mwe-popups-is-tall > div > a > svg{vertical-align:middle}.mwe-popups.mwe-popups-is-tall .mwe-popups-extract{width:215px;height:180px;overflow:hidden; float:left}.mwe-popups.mwe-popups-is-tall footer{width:215px}.mwe-popups.mwe-popups-is-not-tall{width:320px}.mwe-popups.mwe-popups-is-not-tall .mwe-popups-extract{min-height:40px;max-height:140px;overflow:hidden;margin-bottom:47px;padding-bottom:0}.mwe-popups.mwe-popups-is-not-tall footer{width:290px}.mwe-popups.mwe-popups-type-generic .mwe-popups-extract,.mwe-popups.mwe-popups-type-disambiguation .mwe-popups-extract{min-height:auto;padding-top:4px;margin-bottom:60px;margin-top:0}.mwe-popups.mwe-popups-type-generic .mwe-popups-read-link,.mwe-popups.mwe-popups-type-disambiguation .mwe-popups-read-link{font-weight:bold;font-size:12px}.mwe-popups.mwe-popups-type-generic .mwe-popups-extract:hover + footer .mwe-popups-read-link,.mwe-popups.mwe-popups-type-disambiguation .mwe-popups-extract:hover + footer .mwe-popups-read-link{text-decoration:underline}.mwe-popups.mwe-popups-no-image-tri:after{content:'';position:absolute;border:11px solid transparent;border-top:0;border-bottom:11px solid #ffffff;top:-7px; left:7px}.mwe-popups.mwe-popups-no-image-tri:before{content:'';position:absolute;border:8px solid transparent;border-top:0;border-bottom:8px solid #a2a9b1;top:-8px; left:10px}.mwe-popups.flipped_x.mwe-popups-no-image-tri:after{ left:auto; right:7px}.mwe-popups.flipped_x.mwe-popups-no-image-tri:before{ left:auto; right:10px}.mwe-popups.mwe-popups-image-tri:before{z-index:111;content:'';position:absolute;border:9px solid transparent;border-top:0;border-bottom:9px solid #a2a9b1;top:-9px; left:9px}.mwe-popups.mwe-popups-image-tri:after{content:'';position:absolute;border:12px solid transparent;border-top:0;border-bottom:12px solid #ffffff;top:-8px; left:6px;z-index:112}.mwe-popups.mwe-popups-image-tri.flipped_x:before{z-index:111;content:'';position:absolute;border:9px solid transparent;border-top:0;border-bottom:9px solid #a2a9b1;top:-9px; left:273px}.mwe-popups.mwe-popups-image-tri.flipped_x:after{content:'';position:absolute;border:12px solid transparent;border-top:0;border-bottom:12px solid #ffffff;top:-8px; left:269px;z-index:112}.mwe-popups.mwe-popups-image-tri .mwe-popups-extract{padding-top:32px;margin-top:190px}.mwe-popups.mwe-popups-image-tri > div > a > svg{margin-top:-8px;position:absolute;z-index:113; left:0}.mwe-popups.flipped_x.mwe-popups-is-tall{min-height:242px}.mwe-popups.flipped_x.mwe-popups-is-tall:before{z-index:111;content:'';position:absolute;border:9px solid transparent;border-top:0;border-bottom:9px solid #a2a9b1;top:-9px; left:420px}.mwe-popups.flipped_x.mwe-popups-is-tall > div > a > svg{margin:0;margin-top:-8px;margin-bottom:-7px;position:absolute;z-index:113; right:0}.mwe-popups.flipped_x_y:before{z-index:111;content:'';position:absolute;border:9px solid transparent;border-bottom:0;border-top:9px solid #a2a9b1;bottom:-9px; left:272px}.mwe-popups.flipped_x_y:after{content:'';position:absolute;border:12px solid transparent;border-bottom:0;border-top:12px solid #ffffff;bottom:-8px; left:269px;z-index:112}.mwe-popups.flipped_x_y.mwe-popups-is-tall{min-height:242px}.mwe-popups.flipped_x_y.mwe-popups-is-tall:after{z-index:112;content:'';position:absolute;border:12px solid transparent;border-bottom:0;border-top:12px solid #ffffff;bottom:-8px; left:417px}.mwe-popups.flipped_x_y.mwe-popups-is-tall:before{z-index:111;content:'';position:absolute;border:9px solid transparent;border-bottom:0;border-top:9px solid #a2a9b1;bottom:-9px; left:420px}.mwe-popups.flipped_x_y.mwe-popups-is-tall > div > a > svg{position:absolute;z-index:113;margin:0;margin-bottom:-9px; right:0}.mwe-popups.flipped_y:after{content:'';position:absolute;border:11px solid transparent;border-bottom:0;border-top:11px solid #ffffff;bottom:-7px; left:7px}.mwe-popups.flipped_y:before{content:'';position:absolute;border:8px solid transparent;border-bottom:0;border-top:8px solid #a2a9b1;bottom:-8px; left:10px} @-webkit-keyframes mwe-popups-fade-in-up{0%{opacity:0;-webkit-transform:translate(0,20px);-moz-transform:translate(0,20px);-ms-transform:translate(0,20px);transform:translate(0,20px)}100%{opacity:1;-webkit-transform:translate(0,0);-moz-transform:translate(0,0);-ms-transform:translate(0,0);transform:translate(0,0)}}@-moz-keyframes mwe-popups-fade-in-up{0%{opacity:0;-webkit-transform:translate(0,20px);-moz-transform:translate(0,20px);-ms-transform:translate(0,20px);transform:translate(0,20px)}100%{opacity:1;-webkit-transform:translate(0,0);-moz-transform:translate(0,0);-ms-transform:translate(0,0);transform:translate(0,0)}}@keyframes mwe-popups-fade-in-up{0%{opacity:0;-webkit-transform:translate(0,20px);-moz-transform:translate(0,20px);-ms-transform:translate(0,20px);transform:translate(0,20px)}100%{opacity:1;-webkit-transform:translate(0,0);-moz-transform:translate(0,0);-ms-transform:translate(0,0);transform:translate(0,0)}}@-webkit-keyframes mwe-popups-fade-in-down{0%{opacity:0;-webkit-transform:translate(0,-20px);-moz-transform:translate(0,-20px);-ms-transform:translate(0,-20px);transform:translate(0,-20px)}100%{opacity:1;-webkit-transform:translate(0,0);-moz-transform:translate(0,0);-ms-transform:translate(0,0);transform:translate(0,0)}}@-moz-keyframes mwe-popups-fade-in-down{0%{opacity:0;-webkit-transform:translate(0,-20px);-moz-transform:translate(0,-20px);-ms-transform:translate(0,-20px);transform:translate(0,-20px)}100%{opacity:1;-webkit-transform:translate(0,0);-moz-transform:translate(0,0);-ms-transform:translate(0,0);transform:translate(0,0)}}@keyframes mwe-popups-fade-in-down{0%{opacity:0;-webkit-transform:translate(0,-20px);-moz-transform:translate(0,-20px);-ms-transform:translate(0,-20px);transform:translate(0,-20px)}100%{opacity:1;-webkit-transform:translate(0,0);-moz-transform:translate(0,0);-ms-transform:translate(0,0);transform:translate(0,0)}}@-webkit-keyframes mwe-popups-fade-out-down{0%{opacity:1;-webkit-transform:translate(0,0);-moz-transform:translate(0,0);-ms-transform:translate(0,0);transform:translate(0,0)}100%{opacity:0;-webkit-transform:translate(0,20px);-moz-transform:translate(0,20px);-ms-transform:translate(0,20px);transform:translate(0,20px)}}@-moz-keyframes mwe-popups-fade-out-down{0%{opacity:1;-webkit-transform:translate(0,0);-moz-transform:translate(0,0);-ms-transform:translate(0,0);transform:translate(0,0)}100%{opacity:0;-webkit-transform:translate(0,20px);-moz-transform:translate(0,20px);-ms-transform:translate(0,20px);transform:translate(0,20px)}}@keyframes mwe-popups-fade-out-down{0%{opacity:1;-webkit-transform:translate(0,0);-moz-transform:translate(0,0);-ms-transform:translate(0,0);transform:translate(0,0)}100%{opacity:0;-webkit-transform:translate(0,20px);-moz-transform:translate(0,20px);-ms-transform:translate(0,20px);transform:translate(0,20px)}}@-webkit-keyframes mwe-popups-fade-out-up{0%{opacity:1;-webkit-transform:translate(0,0);-moz-transform:translate(0,0);-ms-transform:translate(0,0);transform:translate(0,0)}100%{opacity:0;-webkit-transform:translate(0,-20px);-moz-transform:translate(0,-20px);-ms-transform:translate(0,-20px);transform:translate(0,-20px)}}@-moz-keyframes mwe-popups-fade-out-up{0%{opacity:1;-webkit-transform:translate(0,0);-moz-transform:translate(0,0);-ms-transform:translate(0,0);transform:translate(0,0)}100%{opacity:0;-webkit-transform:translate(0,-20px);-moz-transform:translate(0,-20px);-ms-transform:translate(0,-20px);transform:translate(0,-20px)}}@keyframes mwe-popups-fade-out-up{0%{opacity:1;-webkit-transform:translate(0,0);-moz-transform:translate(0,0);-ms-transform:translate(0,0);transform:translate(0,0)}100%{opacity:0;-webkit-transform:translate(0,-20px);-moz-transform:translate(0,-20px);-ms-transform:translate(0,-20px);transform:translate(0,-20px)}}.mwe-popups-fade-in-up{-webkit-animation:mwe-popups-fade-in-up 0.2s ease forwards;-moz-animation:mwe-popups-fade-in-up 0.2s ease forwards;animation:mwe-popups-fade-in-up 0.2s ease forwards}.mwe-popups-fade-in-down{-webkit-animation:mwe-popups-fade-in-down 0.2s ease forwards;-moz-animation:mwe-popups-fade-in-down 0.2s ease forwards;animation:mwe-popups-fade-in-down 0.2s ease forwards}.mwe-popups-fade-out-down{-webkit-animation:mwe-popups-fade-out-down 0.2s ease forwards;-moz-animation:mwe-popups-fade-out-down 0.2s ease forwards;animation:mwe-popups-fade-out-down 0.2s ease forwards}.mwe-popups-fade-out-up{-webkit-animation:mwe-popups-fade-out-up 0.2s ease forwards;-moz-animation:mwe-popups-fade-out-up 0.2s ease forwards;animation:mwe-popups-fade-out-up 0.2s ease forwards}     #mwe-popups-settings{position:fixed;z-index:1000;background:#fff;width:420px;border:1px solid #a2a9b1;box-shadow:0 2px 2px 0 rgba(0,0,0,0.25);border-radius:2px;font-size:14px}#mwe-popups-settings header{-webkit-box-sizing:border-box;-moz-box-sizing:border-box;box-sizing:border-box;border-bottom:1px solid #c8ccd1;position:relative;display:table;width:100%;padding:5px 7px 5px 0}#mwe-popups-settings header > div{display:table-cell;width:3.5em;vertical-align:middle;cursor:pointer}#mwe-popups-settings header h1{margin-bottom:0.6em;padding-top:0.5em;border:0;width:100%;font-family:sans-serif;font-size:18px;font-weight:bold;text-align:center}#mwe-popups-settings .mwe-ui-icon-popups-close{opacity:0.87;-webkit-transition:opacity 100ms;-moz-transition:opacity 100ms;transition:opacity 100ms}#mwe-popups-settings .mwe-ui-icon-popups-close:hover{opacity:0.73}#mwe-popups-settings .mwe-ui-icon-popups-close:active{opacity:1}#mwe-popups-settings main{display:block;width:350px;padding:32px 0 24px;margin:0 auto}#mwe-popups-settings main p{color:#54595d;font-size:17px;margin:16px 0 0}#mwe-popups-settings main p:first-child{margin-top:0}#mwe-popups-settings main form img,#mwe-popups-settings main form input,#mwe-popups-settings main form label{vertical-align:top}#mwe-popups-settings main form img{margin-right:60px}#mwe-popups-settings main form input{display:inline-block;margin:0 10px 0 0;padding:0}#mwe-popups-settings main form label{font-size:13px;display:inline-block;line-height:16px;width:300px}#mwe-popups-settings main form label > span{color:#000;font-size:18px;font-weight:bold;display:block;margin-bottom:5px;line-height:18px}.mwe-popups-settings-help{background-image:url(/w/extensions/Popups/resources/ext.popups.main/images/footer-ltr.png?3533c);background-image:linear-gradient(transparent,transparent),url("data:image/svg+xml,%3Csvg xmlns=%22http://www.w3.org/2000/svg%22 width=%22230%22 height=%22179%22 xmlns:xlink=%22http://www.w3.org/1999/xlink%22 viewBox=%220 0 230 179%22%3E %3Cdefs%3E %3Crect id=%22a%22 width=%22201%22 height=%2213%22 rx=%222%22/%3E %3Crect id=%22b%22 width=%22201%22 height=%22169%22 y=%2210%22 rx=%222%22/%3E %3Crect id=%22c%22 width=%2230%22 height=%222%22 x=%22135%22 y=%22158%22 rx=%221%22/%3E %3C/defs%3E %3Cg fill=%22none%22 fill-rule=%22evenodd%22%3E %3Cg transform=%22matrix%281 0 0 -1 0 13%29%22%3E %3Cuse fill=%22%23f8f9fa%22 xlink:href=%22%23a%22/%3E %3Crect width=%22199%22 height=%2211%22 x=%221%22 y=%221%22 stroke=%22%23a2a9b1%22 stroke-width=%222%22 rx=%222%22/%3E %3C/g%3E %3Cuse fill=%22%23fff%22 xlink:href=%22%23b%22/%3E %3Crect width=%22199%22 height=%22167%22 x=%221%22 y=%2211%22 stroke=%22%23a2a9b1%22 stroke-width=%222%22 rx=%222%22/%3E %3Cg opacity=%22.4%22 transform=%22translate%2867 35%29%22%3E %3Crect width=%2273%22 height=%222%22 y=%227%22 fill=%22%23c8ccd1%22 rx=%221%22/%3E %3Crect width=%2281%22 height=%222%22 y=%2231%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%2232%22 height=%222%22 y=%2285%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%2273%22 height=%222%22 x=%2235%22 y=%2285%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%2217%22 height=%222%22 y=%2245%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%2217%22 height=%222%22 x=%2291%22 y=%2245%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%2268%22 height=%222%22 x=%2220%22 y=%2245%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%2217%22 height=%222%22 y=%2278%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%2237%22 height=%222%22 x=%2272%22 y=%2278%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%2249%22 height=%222%22 x=%2220%22 y=%2278%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%2224%22 height=%222%22 x=%2284%22 y=%2231%22 fill=%22%2372777d%22 rx=%221%22 transform=%22matrix%28-1 0 0 1 192 0%29%22/%3E %3Crect width=%2281%22 height=%222%22 y=%2266%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%2214%22 height=%222%22 x=%2254%22 y=%2224%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%2237%22 height=%222%22 x=%2271%22 y=%2224%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%2251%22 height=%222%22 y=%2224%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%22108%22 height=%222%22 y=%2259%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%22108%22 height=%222%22 y=%2252%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%22108%22 height=%222%22 y=%2292%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%22108%22 height=%222%22 y=%2238%22 fill=%22%2372777d%22 rx=%221%22/%3E %3Crect width=%2251%22 height=%222%22 fill=%22%2372777d%22 rx=%221%22/%3E %3C/g%3E %3Crect width=%2230%22 height=%222%22 x=%2267%22 y=%22158%22 fill=%22%2372777d%22 opacity=%22.4%22 rx=%221%22/%3E %3Crect width=%2230%22 height=%222%22 x=%2299%22 y=%22158%22 fill=%22%2372777d%22 opacity=%22.4%22 rx=%221%22/%3E %3Cuse fill=%22%2336c%22 xlink:href=%22%23c%22/%3E %3Crect width=%2233%22 height=%225%22 x=%22133.5%22 y=%22156.5%22 stroke=%22%23ffc057%22 stroke-opacity=%22.447%22 stroke-width=%223%22 rx=%222.5%22/%3E %3Ccircle cx=%2234%22 cy=%2249%22 r=%2219%22 fill=%22%23eaecf0%22/%3E %3Cg fill=%22%23a2a9b1%22 transform=%22translate%285 5%29%22%3E %3Ccircle cx=%221.5%22 cy=%221.5%22 r=%221.5%22/%3E %3Ccircle cx=%226%22 cy=%221.5%22 r=%221.5%22/%3E %3Ccircle cx=%2210.5%22 cy=%221.5%22 r=%221.5%22/%3E %3C/g%3E %3Cpath stroke=%22%23ff00af%22 d=%22M174.5 159.5h54.01%22 stroke-linecap=%22square%22/%3E %3C/g%3E %3C/svg%3E");background-position:top left;background-repeat:no-repeat;background-size:180px 140px;font-size:13px;font-weight:800;height:140px;margin:40px;position:relative}.mwe-popups-settings-help p{left:180px;bottom:20px;position:absolute}.mwe-popups-overlay{background-color:rgba(255,255,255,0.9);z-index:999;position:fixed;height:100%;width:100%;top:0;bottom:0;left:0;right:0;display:flex;justify-content:center;align-items:center}</STYLE><STYLE>
.ve-activated .ve-init-mw-desktopArticleTarget-editableContent #toc,.ve-activated #siteNotice,.ve-activated .mw-indicators,.ve-activated #t-print,.ve-activated #t-permalink,.ve-activated #p-coll-print_export,.ve-activated #t-cite,.ve-deactivating .ve-ui-surface,.ve-active .ve-init-mw-desktopArticleTarget-editableContent,.ve-active .ve-init-mw-tempWikitextEditorWidget{display:none} .ve-activating .ve-ui-surface{height:0;padding:0 !important; overflow:hidden} .ve-loading #content > :not(.ve-init-mw-desktopArticleTarget-loading-overlay), .ve-activated .ve-init-mw-desktopArticleTarget-uneditableContent{pointer-events:none;-webkit-user-select:none;-moz-user-select:none;-ms-user-select:none;user-select:none;opacity:0.5}.ve-activated #catlinks{cursor:pointer}.ve-activated #catlinks a{opacity:1} .ve-activated #content{position:relative} .ve-init-mw-desktopArticleTarget-loading-overlay{position:absolute;top:1.25em;left:0;right:0;z-index:1;margin-top:-0.5em}.ve-init-mw-desktopArticleTarget-progress{height:1em;overflow:hidden;margin:0 25%}.ve-init-mw-desktopArticleTarget-progress-bar{height:1em;width:0}.ve-init-mw-desktopArticleTarget-toolbarPlaceholder{transition:height 250ms ease;height:0; } .oo-ui-element-hidden{display:none !important; } .mw-editsection{white-space:nowrap; unicode-bidi:-moz-isolate;unicode-bidi:-webkit-isolate;unicode-bidi:isolate}.mw-editsection-divider{color:#54595d} .ve-init-mw-desktopArticleTarget-progress{height:0.75em;border:1px solid #36c;background:#fff;border-radius:2px;box-shadow:0 0.1em 0 0 rgba(0,0,0,0.15)}.ve-init-mw-desktopArticleTarget-progress-bar{height:0.75em;background:#36c}.ve-init-mw-desktopArticleTarget-toolbar,.ve-init-mw-desktopArticleTarget-toolbarPlaceholder{font-size:0.875em; margin:-1.14em -1.14em 1.14em -1.14em; }@media screen and (min-width:982px){.ve-init-mw-desktopArticleTarget-toolbar,.ve-init-mw-desktopArticleTarget-toolbarPlaceholder{ margin:-1.43em -1.71em 1.43em -1.71em}}.ve-init-mw-desktopArticleTarget-toolbarPlaceholder{border-bottom:1px solid #c8ccd1;box-shadow:0 1px 1px 0 rgba(0,0,0,0.1)}.ve-init-mw-desktopArticleTarget-toolbarPlaceholder-open{height:40px}</STYLE><META name="ResourceLoaderDynamicStyles" content="" />
<LINK rel="stylesheet" href="/w/load.php?debug=false&lang=en&modules=ext.gadget.charinsert-styles&only=styles&skin=vector" />
<LINK rel="stylesheet" href="/w/load.php?debug=false&lang=en&modules=site.styles&only=styles&skin=vector" />
<META name="generator" content="MediaWiki 1.31.0-wmf.28" />
<META name="referrer" content="origin" />
<META name="referrer" content="origin-when-crossorigin" />
<META name="referrer" content="origin-when-cross-origin" />
<LINK rel="alternate" href="android-app://org.wikipedia/http/en.m.wikipedia.org/wiki/Colony_(biology)" />
<LINK rel="alternate" type="application/x-wiki" title="Edit this page" href="/w/index.php?title=Colony_(biology)&action=edit" />
<LINK rel="edit" title="Edit this page" href="/w/index.php?title=Colony_(biology)&action=edit" />
<LINK rel="apple-touch-icon" href="/static/apple-touch/wikipedia.png" />
<LINK rel="shortcut icon" href="/static/favicon/wikipedia.ico" />
<LINK rel="search" type="application/opensearchdescription+xml" href="/w/opensearch_desc.php" title="Wikipedia (en)" />
<LINK rel="EditURI" type="application/rsd+xml" href="//en.wikipedia.org/w/api.php?action=rsd" />
<LINK rel="license" href="//creativecommons.org/licenses/by-sa/3.0/" />
<LINK rel="canonical" href="https://en.wikipedia.org/wiki/Colony_(biology)" />
<LINK rel="dns-prefetch" href="//login.wikimedia.org" />
<LINK rel="dns-prefetch" href="//meta.wikimedia.org" />
<!--[if lt IE 9]><script src="/w/load.php?debug=false&lang=en&modules=html5shiv&only=scripts&skin=vector&sync=1"></script><![endif]-->
</HEAD>
<BODY class="mediawiki ltr sitedir-ltr mw-hide-empty-elt ns-0 ns-subject page-Colony_biology rootpage-Colony_biology skin-vector action-view">		<DIV id="mw-page-base" class="noprint"></DIV>
		<DIV id="mw-head-base" class="noprint"></DIV>
		<DIV id="content" class="mw-body" role="main">
			<A id="top"></A>
			<DIV id="siteNotice" class="mw-body-content"><DIV id="centralNotice"></DIV><!-- CentralNotice --></DIV><DIV class="mw-indicators mw-body-content">
</DIV>
<H1 id="firstHeading" class="firstHeading" lang="en">Colony (biology)</H1>			<DIV id="bodyContent" class="mw-body-content">
				<DIV id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</DIV>				<DIV id="contentSub"></DIV>
								<DIV id="jump-to-nav" class="mw-jump">
					Jump to:					<A href="#mw-head">navigation</A>, 					<A href="#p-search">search</A>
				</DIV>
				<DIV id="mw-content-text" lang="en" dir="ltr" class="mw-content-ltr"><DIV class="mw-parser-output"><P>In <A href="/wiki/Biology" title="Biology">biology</A>, a <B>colony</B> is composed of two or more <A href="/wiki/Conspecific" class="mw-redirect" title="Conspecific">conspecific</A> individuals living in close association with, or connected to, one another. This association is usually for mutual benefit such as stronger defense or the ability to attack bigger prey.<SUP id="cite_ref-Jackson1977_1-0" class="reference"><A href="#cite_note-Jackson1977-1">[1]</A></SUP> It is a cluster of identical cells (clones) on the surface of (or within) a solid medium, usually derived from a single parent cell, as in bacterial colony.<SUP id="cite_ref-2" class="reference"><A href="#cite_note-2">[2]</A></SUP> In contrast, a <A href="/wiki/Solitary_animal" title="Solitary animal">solitary organism</A> is one in which all individuals live independently and have all of the functions needed to survive and reproduce.</P>
<P>Colonies, in the context of development, may be composed of two or more unitary (or solitary) organisms or be <A href="/wiki/Modular" class="mw-redirect" title="Modular">modular</A> organisms. <B>Unitary organisms</B> have determinate development (set life stages) from <A href="/wiki/Zygote" title="Zygote">zygote</A> to adult form and individuals or groups of individuals (colonies) are visually distinct. <B>Modular organisms</B> have indeterminate growth forms (life stages not set) through repeated iteration of genetically identical modules (or individuals), and it can be difficult to distinguish between the colony as a whole and the modules within.<SUP id="cite_ref-3" class="reference"><A href="#cite_note-3">[3]</A></SUP> In the latter case, modules may have specific functions within the colony.</P>
<P>Some organisms are primarily independent and form <A href="/wiki/Facultative" title="Facultative">facultative</A> colonies in reply to environmental conditions while others must live in a colony to survive (<A href="/wiki/Obligate" title="Obligate">obligate</A>). For example, some <A href="/wiki/Carpenter_bee" title="Carpenter bee">carpenter bees</A> will form colonies when a dominant hierarchy is formed between two or more nest foundresses<SUP id="cite_ref-DunnRichards2003_4-0" class="reference"><A href="#cite_note-DunnRichards2003-4">[4]</A></SUP> (facultative colony), while <A href="/wiki/Coral" title="Coral">corals</A> are animals that are physically connected by living tissue (the <A href="/wiki/Coenosarc" title="Coenosarc">coenosarc</A>) that contains a shared <A href="/wiki/Gastrovascular_cavity" title="Gastrovascular cavity">gastrovascular cavity</A>.</P>
<P></P>
<DIV id="toc" class="toc">
<DIV class="toctitle" lang="en" dir="ltr" xml:lang="en">
<H2>Contents</H2>
<SPAN class="toctoggle"> [<A role="button" tabindex="0" class="togglelink">hide</A>] </SPAN></DIV>
<UL>
<LI class="toclevel-1 tocsection-1"><A href="#Colony_types"><SPAN class="tocnumber">1</SPAN> <SPAN class="toctext">Colony types</SPAN></A>
<UL>
<LI class="toclevel-2 tocsection-2"><A href="#Social_colonies"><SPAN class="tocnumber">1.1</SPAN> <SPAN class="toctext">Social colonies</SPAN></A></LI>
<LI class="toclevel-2 tocsection-3"><A href="#Modular_organisms"><SPAN class="tocnumber">1.2</SPAN> <SPAN class="toctext">Modular organisms</SPAN></A></LI>
<LI class="toclevel-2 tocsection-4"><A href="#Microbial_colonies"><SPAN class="tocnumber">1.3</SPAN> <SPAN class="toctext">Microbial colonies</SPAN></A></LI>
</UL>
</LI>
<LI class="toclevel-1 tocsection-5"><A href="#Life_history"><SPAN class="tocnumber">2</SPAN> <SPAN class="toctext">Life history</SPAN></A></LI>
<LI class="toclevel-1 tocsection-6"><A href="#See_also"><SPAN class="tocnumber">3</SPAN> <SPAN class="toctext">See also</SPAN></A></LI>
<LI class="toclevel-1 tocsection-7"><A href="#References"><SPAN class="tocnumber">4</SPAN> <SPAN class="toctext">References</SPAN></A></LI>
</UL>
</DIV>
<P></P>
<H2><SPAN class="mw-headline" id="Colony_types">Colony types</SPAN><SPAN class="mw-editsection"><SPAN class="mw-editsection-bracket">[</SPAN><A href="/w/index.php?title=Colony_(biology)&action=edit&section=1" title="Edit section: Colony types">edit</A><SPAN class="mw-editsection-bracket">]</SPAN></SPAN></H2>
<H3><SPAN class="mw-headline" id="Social_colonies">Social colonies</SPAN><SPAN class="mw-editsection"><SPAN class="mw-editsection-bracket">[</SPAN><A href="/w/index.php?title=Colony_(biology)&action=edit&section=2" title="Edit section: Social colonies">edit</A><SPAN class="mw-editsection-bracket">]</SPAN></SPAN></H3>
<DIV class="thumb tright">
<DIV class="thumbinner" style="width:222px;"><A href="/wiki/File:Colony_Northern_Gannet_Morus_bassanus.jpg" class="image"><IMG alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/0/02/Colony_Northern_Gannet_Morus_bassanus.jpg/220px-Colony_Northern_Gannet_Morus_bassanus.jpg" width="220" height="146" class="thumbimage" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/02/Colony_Northern_Gannet_Morus_bassanus.jpg/330px-Colony_Northern_Gannet_Morus_bassanus.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/02/Colony_Northern_Gannet_Morus_bassanus.jpg/440px-Colony_Northern_Gannet_Morus_bassanus.jpg 2x" data-file-width="1248" data-file-height="830" /></A>
<DIV class="thumbcaption">
<DIV class="magnify"><A href="/wiki/File:Colony_Northern_Gannet_Morus_bassanus.jpg" class="internal" title="Enlarge"></A></DIV>
A breeding colony of <A href="/wiki/Northern_gannet" title="Northern gannet">Northern gannets</A> on the <A href="/wiki/Heligoland" title="Heligoland">Heligoland</A> archipelago in the <A href="/wiki/North_Sea" title="North Sea">North Sea</A>.</DIV>
</DIV>
</DIV>
<P><A href="/wiki/Unicellular" class="mw-redirect" title="Unicellular">Unicellular</A> and <A href="/wiki/Multicellular" class="mw-redirect" title="Multicellular">multicellular</A> unitary organisms may aggregate to form colonies. For example,</P>
<UL>
<LI><A href="/wiki/Protist" title="Protist">Protists</A> such as <A href="/wiki/Slime_mold" title="Slime mold">slime molds</A> are many unicellular organisms that aggregate to form colonies when food resources are hard to come by, as together they are more reactive to chemical cues released by preferred prey.</LI>
<LI><A href="/wiki/Eusociality" title="Eusociality">Eusocial</A> insects like <A href="/wiki/Ant" title="Ant">ants</A> and <A href="/wiki/Honey_bee" title="Honey bee">honey bees</A> are multicellular animals that live in colonies with a highly organized social structure. Colonies of some social insects may be deemed <A href="/wiki/Superorganism" title="Superorganism">superorganisms</A>.</LI>
<LI>Animals such as <A href="/wiki/Human" title="Human">humans</A> and <A href="/wiki/Rodent" title="Rodent">rodents</A> form breeding or <A href="/wiki/Bird_colony" title="Bird colony">nesting colonies</A>, potentially for more successful mating and to better protect offspring.
<UL>
<LI>The <A href="/wiki/Bracken_Cave" title="Bracken Cave">Bracken Cave</A> is the summer home to a colony of around 20 million <A href="/wiki/Mexican_free-tailed_bat" title="Mexican free-tailed bat">Mexican free-tailed bats</A> making it the largest known concentration of mammals.<SUP id="cite_ref-natgeo_5-0" class="reference"><A href="#cite_note-natgeo-5">[5]</A></SUP></LI>
</UL>
</LI>
</UL>
<H3><SPAN class="mw-headline" id="Modular_organisms">Modular organisms</SPAN><SPAN class="mw-editsection"><SPAN class="mw-editsection-bracket">[</SPAN><A href="/w/index.php?title=Colony_(biology)&action=edit&section=3" title="Edit section: Modular organisms">edit</A><SPAN class="mw-editsection-bracket">]</SPAN></SPAN></H3>
<P>Modular organisms are those in which a genet (or genetic individual formed from a <A href="/wiki/Sexual_reproduction" title="Sexual reproduction">sexually-produced</A> <A href="/wiki/Zygote" title="Zygote">zygote</A>) asexually reproduces to form genetically identical clones called <A href="/wiki/Ramet" class="mw-redirect" title="Ramet">ramets</A>.<SUP id="cite_ref-Winston2010_6-0" class="reference"><A href="#cite_note-Winston2010-6">[6]</A></SUP></P>
<P>A <B><A href="/wiki/Clonal_colony" title="Clonal colony">clonal colony</A></B> is when the ramets of a genet live in close proximity or are physically connected. Ramets may have all of the functions needed to survive on their own or be interdependent on other ramets. For example, some <A href="/wiki/Sea_anemone" title="Sea anemone">sea anemones</A> go through the process of pedal laceration in which a genetically identical individual is asexually produced from tissue broken off from the anemone's pedal disc. In plants, clonal colonies are created through the propagation of genetically identical trees by <A href="/wiki/Stolon" title="Stolon">stolons</A> or <A href="/wiki/Rhizome" title="Rhizome">rhizomes</A>.</P>
<P><B>Colonial organisms</B> are <A href="/wiki/Clonal_colonies" class="mw-redirect" title="Clonal colonies">clonal colonies</A> composed of many physically connected, interdependent individuals. Colonial organisms can be unicellular, as in the alga <I><A href="/wiki/Volvox" title="Volvox">Volvox</A></I> (a <A href="/wiki/Coenobium_(morphology)" title="Coenobium (morphology)">coenobium</A>), or <A href="/wiki/Multicellular_organism" title="Multicellular organism">multicellular</A>, as in the <A href="/wiki/Phylum" title="Phylum">phylum</A> <A href="/wiki/Bryozoa" title="Bryozoa">Bryozoa</A>. The former type may have been the first step toward multicellular <A href="/wiki/Organism" title="Organism">organisms</A>.<SUP id="cite_ref-7" class="reference"><A href="#cite_note-7">[7]</A></SUP> Individuals within a multicellular colonial organism may be called <A href="/wiki/Ramet" class="mw-redirect" title="Ramet">ramets</A>, <A href="/wiki/Modularity_(biology)" title="Modularity (biology)">modules</A>, or <A href="/wiki/Zooid" title="Zooid">zooids</A>. Structural and functional variation (<A href="/wiki/Polymorphism_(biology)" title="Polymorphism (biology)">polymorphism</A>), when present, designates ramet responsibilities such as feeding, reproduction, and defense. To that end, being physically connected allows the colonial organism to distribute nutrients and energy obtained by feeding zooids throughout the colony. An example of colonial organisms that is well known are hydrozoans, like Portuguese man o' wars.<SUP id="cite_ref-8" class="reference"><A href="#cite_note-8">[8]</A></SUP></P>
<H3><SPAN class="mw-headline" id="Microbial_colonies">Microbial colonies</SPAN><SPAN class="mw-editsection"><SPAN class="mw-editsection-bracket">[</SPAN><A href="/w/index.php?title=Colony_(biology)&action=edit&section=4" title="Edit section: Microbial colonies">edit</A><SPAN class="mw-editsection-bracket">]</SPAN></SPAN></H3>
<P>A <B>microbial colony</B> is defined as a visible cluster of <A href="/wiki/Microorganism" title="Microorganism">microorganisms</A> growing on the surface of or within a solid medium, presumably cultured from a single cell.<SUP id="cite_ref-9" class="reference"><A href="#cite_note-9">[9]</A></SUP> Because the colony is <A href="/wiki/Clone_(cell_biology)" title="Clone (cell biology)">clonal</A>, with all organisms in it descending from a single ancestor (assuming no <A href="/wiki/Contamination" title="Contamination">contamination</A>), they are genetically identical, except for any <A href="/wiki/Mutation" title="Mutation">mutations</A> (which occur at low frequencies). Obtaining such genetically identical organisms (or pure <A href="/wiki/Strain_(biology)" title="Strain (biology)">strains</A>) can be useful; this is done by spreading organisms on a culture plate and starting a new stock from a single resulting colony.</P>
<P>A <B><A href="/wiki/Biofilm" title="Biofilm">biofilm</A></B> is a colony of <A href="/wiki/Microorganism" title="Microorganism">microorganisms</A> often comprising several species, with properties and capabilities greater than the aggregate of capabilities of the individual organisms.</P>
<H2><SPAN class="mw-headline" id="Life_history">Life history</SPAN><SPAN class="mw-editsection"><SPAN class="mw-editsection-bracket">[</SPAN><A href="/w/index.php?title=Colony_(biology)&action=edit&section=5" title="Edit section: Life history">edit</A><SPAN class="mw-editsection-bracket">]</SPAN></SPAN></H2>
<P>Individuals in social colonies and modular organisms receive benefit to such a lifestyle. For example, it may be easier to seek out food, defend a nesting site, or increase competitive ability against other species. Modular organisms' ability to reproduce asexually in addition to sexually allows them unique benefits that social colonies do not have.<SUP id="cite_ref-Winston2010_6-1" class="reference"><A href="#cite_note-Winston2010-6">[6]</A></SUP></P>
<P>The energy required for sexual reproduction varies based on the frequency and length of reproductive activity, number and size of offspring, and parental care.<SUP id="cite_ref-KunzOrrell_10-0" class="reference"><A href="#cite_note-KunzOrrell-10">[10]</A></SUP> While solitary individuals bear all of those energy costs, individuals in some social colonies share a portion of those costs.</P>
<P>Modular organisms save energy by using asexual reproduction during their life. Energy reserved in this way allows them to put more energy towards colony growth, regenerating lost modules (due to predation or other cause of death), or response to environmental conditions.</P>
<H2><SPAN class="mw-headline" id="See_also">See also</SPAN><SPAN class="mw-editsection"><SPAN class="mw-editsection-bracket">[</SPAN><A href="/w/index.php?title=Colony_(biology)&action=edit&section=6" title="Edit section: See also">edit</A><SPAN class="mw-editsection-bracket">]</SPAN></SPAN></H2>
<TABLE role="presentation" class="mbox-small plainlinks sistersitebox" style="background-color:#f9f9f9;border:1px solid #aaa;color:#000">
<TBODY><TR>
<TD class="mbox-image"><IMG alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/99/Wiktionary-logo-en-v2.svg/40px-Wiktionary-logo-en-v2.svg.png" width="40" height="40" class="noviewer" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/99/Wiktionary-logo-en-v2.svg/60px-Wiktionary-logo-en-v2.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/99/Wiktionary-logo-en-v2.svg/80px-Wiktionary-logo-en-v2.svg.png 2x" data-file-width="391" data-file-height="391" /></TD>
<TD class="mbox-text plainlist">Look up <I><B><A href="https://en.wiktionary.org/wiki/colony" class="extiw" title="wiktionary:colony">colony</A></B></I> in Wiktionary, the free dictionary.</TD>
</TR>
</TBODY></TABLE>
<UL>
<LI><A href="/wiki/Ant_colony" title="Ant colony">Ant colony</A></LI>
<LI><A href="/wiki/Beehive_(beekeeping)" class="mw-redirect" title="Beehive (beekeeping)">Beehive (beekeeping)</A></LI>
<LI><A href="/wiki/Bird_colony" title="Bird colony">Bird colony</A></LI>
<LI><A href="/wiki/Clonal_colony" title="Clonal colony">Clonal colony</A></LI>
<LI><A href="/wiki/Colonisation_(biology)" title="Colonisation (biology)">Colonisation (biology)</A></LI>
<LI><A href="/wiki/Coral_reef" title="Coral reef">Coral reef</A></LI>
<LI><A href="/wiki/Eusociality" title="Eusociality">Eusociality</A></LI>
<LI><A href="/wiki/Superorganism" title="Superorganism">Superorganism</A></LI>
<LI><A href="/wiki/Swarm" class="mw-redirect" title="Swarm">Swarm</A></LI>
<LI><A href="/wiki/Litter_(animal)" title="Litter (animal)">Birth colony</A></LI>
<LI><I><A href="/wiki/Austroplatypus_incompertus" title="Austroplatypus incompertus">Austroplatypus incompertus</A></I></LI>
</UL>
<H2><SPAN class="mw-headline" id="References">References</SPAN><SPAN class="mw-editsection"><SPAN class="mw-editsection-bracket">[</SPAN><A href="/w/index.php?title=Colony_(biology)&action=edit&section=7" title="Edit section: References">edit</A><SPAN class="mw-editsection-bracket">]</SPAN></SPAN></H2>
<DIV class="mw-references-wrap">
<OL class="references">
<LI id="cite_note-Jackson1977-1"><SPAN class="mw-cite-backlink"><B><A href="#cite_ref-Jackson1977_1-0"><SPAN class="cite-accessibility-label">Jump up </SPAN>^</A></B></SPAN> <SPAN class="reference-text"><CITE class="citation journal">Jackson, J.B.C. (1977). "Competition on Marine Hard Substrata: The Adaptive Significance of Solitary and Colonial Strategies". <I>The American Naturalist</I>. <B>111</B> (980): 743–767. <A href="/wiki/Digital_object_identifier" title="Digital object identifier">doi</A>:<A rel="nofollow" class="external text" href="//doi.org/10.1086%2F283203">10.1086/283203</A>.</CITE><SPAN title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+American+Naturalist&rft.atitle=Competition+on+Marine+Hard+Substrata%3A+The+Adaptive+Significance+of+Solitary+and+Colonial+Strategies&rft.volume=111&rft.issue=980&rft.pages=743-767&rft.date=1977&rft_id=info%3Adoi%2F10.1086%2F283203&rft.aulast=Jackson&rft.aufirst=J.B.C.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AColony+%28biology%29" class="Z3988"><SPAN style="display:none;"> </SPAN></SPAN></SPAN></LI>
<LI id="cite_note-2"><SPAN class="mw-cite-backlink"><B><A href="#cite_ref-2"><SPAN class="cite-accessibility-label">Jump up </SPAN>^</A></B></SPAN> <SPAN class="reference-text"><CITE class="citation web"><A rel="nofollow" class="external text" href="http://www.biology-online.org/dictionary/Colony">"Colony – Biology-Online Dictionary"</A>. <I>www.biology-online.org</I><SPAN class="reference-accessdate">. Retrieved <SPAN class="nowrap">2017-05-06</SPAN></SPAN>.</CITE><SPAN title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=www.biology-online.org&rft.atitle=Colony+%E2%80%93+Biology-Online+Dictionary&rft_id=http%3A%2F%2Fwww.biology-online.org%2Fdictionary%2FColony&rfr_id=info%3Asid%2Fen.wikipedia.org%3AColony+%28biology%29" class="Z3988"><SPAN style="display:none;"> </SPAN></SPAN></SPAN></LI>
<LI id="cite_note-3"><SPAN class="mw-cite-backlink"><B><A href="#cite_ref-3"><SPAN class="cite-accessibility-label">Jump up </SPAN>^</A></B></SPAN> <SPAN class="reference-text"><CITE class="citation book">Begon, Michael; et al. (2014). <I>Essentials of Ecology</I> (4th ed.). Wiley. <A href="/wiki/International_Standard_Book_Number" title="International Standard Book Number">ISBN</A> <A href="/wiki/Special:BookSources/0-470-90913-7" title="Special:BookSources/0-470-90913-7">0-470-90913-7</A>.</CITE><SPAN title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Essentials+of+Ecology&rft.edition=4th&rft.pub=Wiley&rft.date=2014&rft.isbn=0-470-90913-7&rft.aulast=Begon&rft.aufirst=Michael&rft.au=Howarth%2C+Robert+W.&rft.au=Townsend%2C+Colin+R.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AColony+%28biology%29" class="Z3988"><SPAN style="display:none;"> </SPAN></SPAN></SPAN></LI>
<LI id="cite_note-DunnRichards2003-4"><SPAN class="mw-cite-backlink"><B><A href="#cite_ref-DunnRichards2003_4-0"><SPAN class="cite-accessibility-label">Jump up </SPAN>^</A></B></SPAN> <SPAN class="reference-text"><CITE class="citation journal">Dunn, T.; Richards, M.H. (2003). "When to bee social: interactions among environmental constraints, incentives, guarding, and relatedness in a facultatively social carpenter bee". <I>Behavioral Ecology</I>. <B>14</B> (3): 417–424. <A href="/wiki/Digital_object_identifier" title="Digital object identifier">doi</A>:<A rel="nofollow" class="external text" href="//doi.org/10.1093%2Fbeheco%2F14.3.417">10.1093/beheco/14.3.417</A>.</CITE><SPAN title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Behavioral+Ecology&rft.atitle=When+to+bee+social%3A+interactions+among+environmental+constraints%2C+incentives%2C+guarding%2C+and+relatedness+in+a+facultatively+social+carpenter+bee&rft.volume=14&rft.issue=3&rft.pages=417-424&rft.date=2003&rft_id=info%3Adoi%2F10.1093%2Fbeheco%2F14.3.417&rft.aulast=Dunn&rft.aufirst=T.&rft.au=Richards%2C+M.H.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AColony+%28biology%29" class="Z3988"><SPAN style="display:none;"> </SPAN></SPAN></SPAN></LI>
<LI id="cite_note-natgeo-5"><SPAN class="mw-cite-backlink"><B><A href="#cite_ref-natgeo_5-0"><SPAN class="cite-accessibility-label">Jump up </SPAN>^</A></B></SPAN> <SPAN class="reference-text"><CITE class="citation journal">Grove, Noel (December 1988). "Quietly Conserving Nature". <I><A href="/wiki/National_Geographic_(magazine)" class="mw-redirect" title="National Geographic (magazine)">National Geographic</A></I>. <B>174</B> (6): 822.</CITE><SPAN title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=National+Geographic&rft.atitle=Quietly+Conserving+Nature&rft.volume=174&rft.issue=6&rft.pages=822&rft.date=1988-12&rft.aulast=Grove&rft.aufirst=Noel&rfr_id=info%3Asid%2Fen.wikipedia.org%3AColony+%28biology%29" class="Z3988"><SPAN style="display:none;"> </SPAN></SPAN></SPAN></LI>
<LI id="cite_note-Winston2010-6"><SPAN class="mw-cite-backlink">^ <A href="#cite_ref-Winston2010_6-0"><SPAN class="cite-accessibility-label">Jump up to: </SPAN><SUP><I><B>a</B></I></SUP></A> <A href="#cite_ref-Winston2010_6-1"><SUP><I><B>b</B></I></SUP></A></SPAN> <SPAN class="reference-text"><CITE class="citation journal">Winston, J. (2010). "Life in the Colonies: Learning the Alien Ways of Colonial Organisms". <I>Integrative and Comparative Biology</I>. <B>50</B> (6): 919–933. <A href="/wiki/Digital_object_identifier" title="Digital object identifier">doi</A>:<A rel="nofollow" class="external text" href="//doi.org/10.1093%2Ficb%2Ficq146">10.1093/icb/icq146</A>. <A href="/wiki/PubMed_Identifier" class="mw-redirect" title="PubMed Identifier">PMID</A> <A rel="nofollow" class="external text" href="//www.ncbi.nlm.nih.gov/pubmed/21714171">21714171</A>.</CITE><SPAN title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Integrative+and+Comparative+Biology&rft.atitle=Life+in+the+Colonies%3A+Learning+the+Alien+Ways+of+Colonial+Organisms&rft.volume=50&rft.issue=6&rft.pages=919-933&rft.date=2010&rft_id=info%3Adoi%2F10.1093%2Ficb%2Ficq146&rft_id=info%3Apmid%2F21714171&rft.aulast=Winston&rft.aufirst=J.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AColony+%28biology%29" class="Z3988"><SPAN style="display:none;"> </SPAN></SPAN></SPAN></LI>
<LI id="cite_note-7"><SPAN class="mw-cite-backlink"><B><A href="#cite_ref-7"><SPAN class="cite-accessibility-label">Jump up </SPAN>^</A></B></SPAN> <SPAN class="reference-text"><CITE class="citation book">Alberts, Bruce; et al. (1994). <A rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/books/NBK28332/"><I>Molecular Biology of the Cell</I></A> (3rd ed.). New York: Garland Science. <A href="/wiki/International_Standard_Book_Number" title="International Standard Book Number">ISBN</A> <A href="/wiki/Special:BookSources/0-8153-1620-8" title="Special:BookSources/0-8153-1620-8">0-8153-1620-8</A><SPAN class="reference-accessdate">. Retrieved <SPAN class="nowrap">2014-06-11</SPAN></SPAN>.</CITE><SPAN title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Molecular+Biology+of+the+Cell&rft.place=New+York&rft.edition=3rd&rft.pub=Garland+Science&rft.date=1994&rft.isbn=0-8153-1620-8&rft.aulast=Alberts&rft.aufirst=Bruce&rft.au=Bray%2C+Dennis&rft.au=Lewis%2C+Julian&rft.au=Raff%2C+Martin&rft.au=Roberts%2C+Keith&rft.au=Watson%2C+James+D.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fbooks%2FNBK28332%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3AColony+%28biology%29" class="Z3988"><SPAN style="display:none;"> </SPAN></SPAN></SPAN></LI>
<LI id="cite_note-8"><SPAN class="mw-cite-backlink"><B><A href="#cite_ref-8"><SPAN class="cite-accessibility-label">Jump up </SPAN>^</A></B></SPAN> <SPAN class="reference-text"><CITE class="citation web"><A rel="nofollow" class="external text" href="http://animaldiversity.org/accounts/Hydrozoa/">"Hydrozoa"</A>. <I>Animal Diversity Web</I><SPAN class="reference-accessdate">. Retrieved <SPAN class="nowrap">2017-05-06</SPAN></SPAN>.</CITE><SPAN title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Animal+Diversity+Web&rft.atitle=Hydrozoa&rft_id=http%3A%2F%2Fanimaldiversity.org%2Faccounts%2FHydrozoa%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3AColony+%28biology%29" class="Z3988"><SPAN style="display:none;"> </SPAN></SPAN></SPAN></LI>
<LI id="cite_note-9"><SPAN class="mw-cite-backlink"><B><A href="#cite_ref-9"><SPAN class="cite-accessibility-label">Jump up </SPAN>^</A></B></SPAN> <SPAN class="reference-text"><CITE class="citation book">Tortora, Gerard J.; Berdell R., Funke; Christine L., Case (2009). <I><A href="/w/index.php?title=Microbiology,_An_Introduction&action=edit&redlink=1" class="new" title="Microbiology, An Introduction (page does not exist)">Microbiology, An Introduction</A></I>. Berlin: <A href="/wiki/Benjamin_Cummings" title="Benjamin Cummings">Benjamin Cummings</A>. pp. 170–171. <A href="/wiki/International_Standard_Book_Number" title="International Standard Book Number">ISBN</A> <A href="/wiki/Special:BookSources/0-321-58420-1" title="Special:BookSources/0-321-58420-1">0-321-58420-1</A>.</CITE><SPAN title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Microbiology%2C+An+Introduction&rft.place=Berlin&rft.pages=170-171&rft.pub=Benjamin+Cummings&rft.date=2009&rft.isbn=0-321-58420-1&rft.aulast=Tortora&rft.aufirst=Gerard+J.&rft.au=Berdell+R.%2C+Funke&rft.au=Christine+L.%2C+Case&rfr_id=info%3Asid%2Fen.wikipedia.org%3AColony+%28biology%29" class="Z3988"><SPAN style="display:none;"> </SPAN></SPAN></SPAN></LI>
<LI id="cite_note-KunzOrrell-10"><SPAN class="mw-cite-backlink"><B><A href="#cite_ref-KunzOrrell_10-0"><SPAN class="cite-accessibility-label">Jump up </SPAN>^</A></B></SPAN> <SPAN class="reference-text"><CITE class="citation journal">Kunz, T.H.; Orrell, K.S. (2004). "Energy Costs of Reproduction". <I>Encyclopedia of Energy</I>. <B>5</B>: 423–442.</CITE><SPAN title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Encyclopedia+of+Energy&rft.atitle=Energy+Costs+of+Reproduction&rft.volume=5&rft.pages=423-442&rft.date=2004&rft.aulast=Kunz&rft.aufirst=T.H.&rft.au=Orrell%2C+K.S.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AColony+%28biology%29" class="Z3988"><SPAN style="display:none;"> </SPAN></SPAN></SPAN></LI>
</OL>
</DIV>

<!--
NewPP limit report
Parsed by mw1278
Cached time: 20180408211521
Cache expiry: 1900800
Dynamic content: false
CPU time usage: 0.112 seconds
Real time usage: 0.141 seconds
Preprocessor visited node count: 574/1000000
Preprocessor generated node count: 0/1500000
Post‐expand include size: 16116/2097152 bytes
Template argument size: 104/2097152 bytes
Highest expansion depth: 8/40
Expensive parser function count: 0/500
Unstrip recursion depth: 0/20
Unstrip post‐expand size: 10785/5000000 bytes
Lua time usage: 0.047/10.000 seconds
Lua memory usage: 2.51 MB/50 MB
-->
<!--
Transclusion expansion time report (%,ms,calls,template)
100.00%   92.695      1 -total
 49.65%   46.020      5 Template:Cite_journal
 25.21%   23.367      1 Template:Wiktionary
 23.37%   21.659      1 Template:Sister_project
 21.00%   19.465      1 Template:Side_box
 11.69%   10.839      3 Template:Cite_book
  8.87%    8.226      2 Template:Cite_web
-->
</DIV>
<!-- Saved in parser cache with key enwiki:pcache:idhash:161296-0!canonical and timestamp 20180408211521 and revision id 835462497
 -->
<NOSCRIPT><img src="//en.wikipedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" title="" width="1" height="1" style="border: none; position: absolute;" /></NOSCRIPT></DIV>					<DIV class="printfooter">
						Retrieved from "<A dir="ltr" href="https://en.wikipedia.org/w/index.php?title=Colony_(biology)&oldid=835462497">https://en.wikipedia.org/w/index.php?title=Colony_(biology)&oldid=835462497</A>"					</DIV>
				<DIV id="catlinks" class="catlinks" data-mw="interface"><DIV id="mw-normal-catlinks" class="mw-normal-catlinks"><A href="/wiki/Help:Category" title="Help:Category">Categories</A>: <UL><LI><A href="/wiki/Category:Community_ecology" title="Category:Community ecology">Community ecology</A></LI><LI><A href="/wiki/Category:Microbiology_terms" title="Category:Microbiology terms">Microbiology terms</A></LI><LI><A href="/wiki/Category:Habitat" title="Category:Habitat">Habitat</A></LI><LI><A href="/wiki/Category:Environmental_terminology" title="Category:Environmental terminology">Environmental terminology</A></LI></UL></DIV></DIV>				<DIV class="visualClear"></DIV>
							</DIV>
		</DIV>
		<DIV id="mw-navigation">
			<H2>Navigation menu</H2>
			<DIV id="mw-head">
									<DIV id="p-personal" role="navigation" class="" aria-labelledby="p-personal-label">
						<H3 id="p-personal-label">Personal tools</H3>
						<UL>
							<LI id="pt-anonuserpage">Not logged in</LI><LI id="pt-anontalk"><A href="/wiki/Special:MyTalk" title="Discussion about edits from this IP address [ctrl-alt-n]" accesskey="n">Talk</A></LI><LI id="pt-anoncontribs"><A href="/wiki/Special:MyContributions" title="A list of edits made from this IP address [ctrl-alt-y]" accesskey="y">Contributions</A></LI><LI id="pt-createaccount"><A href="/w/index.php?title=Special:CreateAccount&returnto=Colony+%28biology%29" title="You are encouraged to create an account and log in; however, it is not mandatory">Create account</A></LI><LI id="pt-login"><A href="/w/index.php?title=Special:UserLogin&returnto=Colony+%28biology%29" title="You're encouraged to log in; however, it's not mandatory. [ctrl-alt-o]" accesskey="o">Log in</A></LI>						</UL>
					</DIV>
									<DIV id="left-navigation">
										<DIV id="p-namespaces" role="navigation" class="vectorTabs" aria-labelledby="p-namespaces-label">
						<H3 id="p-namespaces-label">Namespaces</H3>
						<UL>
							<LI id="ca-nstab-main" class="selected"><SPAN><A href="/wiki/Colony_(biology)" title="View the content page [ctrl-alt-c]" accesskey="c">Article</A></SPAN></LI><LI id="ca-talk"><SPAN><A href="/wiki/Talk:Colony_(biology)" rel="discussion" title="Discussion about the content page [ctrl-alt-t]" accesskey="t">Talk</A></SPAN></LI>						</UL>
					</DIV>
										<DIV id="p-variants" role="navigation" class="vectorMenu emptyPortlet" aria-labelledby="p-variants-label">
												<INPUT type="checkbox" class="vectorMenuCheckbox" aria-labelledby="p-variants-label" />
						<H3 id="p-variants-label">
							<SPAN>Variants</SPAN>
						</H3>
						<DIV class="menu">
							<UL>
															</UL>
						</DIV>
					</DIV>
									</DIV>
				<DIV id="right-navigation">
										<DIV id="p-views" role="navigation" class="vectorTabs" aria-labelledby="p-views-label">
						<H3 id="p-views-label">Views</H3>
						<UL>
							<LI id="ca-view" class="collapsible selected"><SPAN><A href="/wiki/Colony_(biology)">Read</A></SPAN></LI><LI id="ca-edit" class="collapsible"><SPAN><A href="/w/index.php?title=Colony_(biology)&action=edit" title="Edit this page [ctrl-alt-e]" accesskey="e">Edit</A></SPAN></LI><LI id="ca-history" class="collapsible"><SPAN><A href="/w/index.php?title=Colony_(biology)&action=history" title="Past revisions of this page [ctrl-alt-h]" accesskey="h">View history</A></SPAN></LI>						</UL>
					</DIV>
										<DIV id="p-cactions" role="navigation" class="vectorMenu emptyPortlet" aria-labelledby="p-cactions-label">
						<INPUT type="checkbox" class="vectorMenuCheckbox" aria-labelledby="p-cactions-label" />
						<H3 id="p-cactions-label"><SPAN>More</SPAN></H3>
						<DIV class="menu">
							<UL>
															</UL>
						</DIV>
					</DIV>
										<DIV id="p-search" role="search">
						<H3>
							<LABEL for="searchInput">Search</LABEL>
						</H3>
						<FORM action="/w/index.php" id="searchform">
							<DIV id="simpleSearch">
								<INPUT type="search" name="search" placeholder="Search Wikipedia" title="Search Wikipedia [ctrl-alt-f]" accesskey="f" id="searchInput" tabindex="1" autocomplete="off" /><INPUT type="hidden" value="Special:Search" name="title" /><INPUT type="submit" name="go" value="Go" title="Go to a page with this exact name if it exists" id="searchButton" class="searchButton" />							</DIV>
						</FORM>
					</DIV>
									</DIV>
			</DIV>
			<DIV id="mw-panel">
				<DIV id="p-logo" role="banner"><A class="mw-wiki-logo" href="/wiki/Main_Page" title="Visit the main page"></A></DIV>
						<DIV class="portal" role="navigation" id="p-navigation" aria-labelledby="p-navigation-label">
			<H3 id="p-navigation-label">Navigation</H3>
			<DIV class="body">
								<UL>
					<LI id="n-mainpage-description"><A href="/wiki/Main_Page" title="Visit the main page [ctrl-alt-z]" accesskey="z">Main page</A></LI><LI id="n-contents"><A href="/wiki/Portal:Contents" title="Guides to browsing Wikipedia">Contents</A></LI><LI id="n-featuredcontent"><A href="/wiki/Portal:Featured_content" title="Featured content – the best of Wikipedia">Featured content</A></LI><LI id="n-currentevents"><A href="/wiki/Portal:Current_events" title="Find background information on current events">Current events</A></LI><LI id="n-randompage"><A href="/wiki/Special:Random" title="Load a random article [ctrl-alt-x]" accesskey="x">Random article</A></LI><LI id="n-sitesupport"><A href="https://donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&utm_medium=sidebar&utm_campaign=C13_en.wikipedia.org&uselang=en" title="Support us">Donate to Wikipedia</A></LI><LI id="n-shoplink"><A href="//shop.wikimedia.org" title="Visit the Wikipedia store">Wikipedia store</A></LI>				</UL>
							</DIV>
		</DIV>
			<DIV class="portal" role="navigation" id="p-interaction" aria-labelledby="p-interaction-label">
			<H3 id="p-interaction-label">Interaction</H3>
			<DIV class="body">
								<UL>
					<LI id="n-help"><A href="/wiki/Help:Contents" title="Guidance on how to use and edit Wikipedia">Help</A></LI><LI id="n-aboutsite"><A href="/wiki/Wikipedia:About" title="Find out about Wikipedia">About Wikipedia</A></LI><LI id="n-portal"><A href="/wiki/Wikipedia:Community_portal" title="About the project, what you can do, where to find things">Community portal</A></LI><LI id="n-recentchanges"><A href="/wiki/Special:RecentChanges" title="A list of recent changes in the wiki [ctrl-alt-r]" accesskey="r">Recent changes</A></LI><LI id="n-contactpage"><A href="//en.wikipedia.org/wiki/Wikipedia:Contact_us" title="How to contact Wikipedia">Contact page</A></LI>				</UL>
							</DIV>
		</DIV>
			<DIV class="portal" role="navigation" id="p-tb" aria-labelledby="p-tb-label">
			<H3 id="p-tb-label">Tools</H3>
			<DIV class="body">
								<UL>
					<LI id="t-whatlinkshere"><A href="/wiki/Special:WhatLinksHere/Colony_(biology)" title="List of all English Wikipedia pages containing links to this page [ctrl-alt-j]" accesskey="j">What links here</A></LI><LI id="t-recentchangeslinked"><A href="/wiki/Special:RecentChangesLinked/Colony_(biology)" rel="nofollow" title="Recent changes in pages linked from this page [ctrl-alt-k]" accesskey="k">Related changes</A></LI><LI id="t-upload"><A href="/wiki/Wikipedia:File_Upload_Wizard" title="Upload files [ctrl-alt-u]" accesskey="u">Upload file</A></LI><LI id="t-specialpages"><A href="/wiki/Special:SpecialPages" title="A list of all special pages [ctrl-alt-q]" accesskey="q">Special pages</A></LI><LI id="t-permalink"><A href="/w/index.php?title=Colony_(biology)&oldid=835462497" title="Permanent link to this revision of the page">Permanent link</A></LI><LI id="t-info"><A href="/w/index.php?title=Colony_(biology)&action=info" title="More information about this page">Page information</A></LI><LI id="t-wikibase"><A href="https://www.wikidata.org/wiki/Special:EntityPage/Q354341" title="Link to connected data repository item [ctrl-alt-g]" accesskey="g">Wikidata item</A></LI><LI id="t-cite"><A href="/w/index.php?title=Special:CiteThisPage&page=Colony_%28biology%29&id=835462497" title="Information on how to cite this page">Cite this page</A></LI>				</UL>
							</DIV>
		</DIV>
			<DIV class="portal" role="navigation" id="p-coll-print_export" aria-labelledby="p-coll-print_export-label">
			<H3 id="p-coll-print_export-label">Print/export</H3>
			<DIV class="body">
								<UL>
					<LI id="coll-create_a_book"><A href="/w/index.php?title=Special:Book&bookcmd=book_creator&referer=Colony+%28biology%29">Create a book</A></LI><LI id="coll-download-as-rdf2latex"><A href="/w/index.php?title=Special:ElectronPdf&page=Colony+%28biology%29&action=show-download-screen">Download as PDF</A></LI><LI id="t-print"><A href="/w/index.php?title=Colony_(biology)&printable=yes" title="Printable version of this page [ctrl-alt-p]" accesskey="p">Printable version</A></LI>				</UL>
							</DIV>
		</DIV>
			<DIV class="portal" role="navigation" id="p-lang" aria-labelledby="p-lang-label"><BUTTON class="uls-settings-trigger" title="Language settings"></BUTTON>
			<H3 id="p-lang-label">Languages</H3>
			<DIV class="body">
								<UL>
					<LI class="interlanguage-link interwiki-ar"><A href="https://ar.wikipedia.org/wiki/%D9%85%D8%B3%D8%AA%D8%B9%D9%85%D8%B1%D8%A9_(%D8%A3%D8%AD%D9%8A%D8%A7%D8%A1)" title="مستعمرة (أحياء) – Arabic" lang="ar" hreflang="ar" class="interlanguage-link-target">العربية</A></LI><LI class="interlanguage-link interwiki-az" style="display: none;"><A href="https://az.wikipedia.org/wiki/Orqanizml%C9%99rin_koloniyas%C4%B1" title="Orqanizmlərin koloniyası – Azerbaijani" lang="az" hreflang="az" class="interlanguage-link-target">Azərbaycanca</A></LI><LI class="interlanguage-link interwiki-bh" style="display: none;"><A href="https://bh.wikipedia.org/wiki/%E0%A4%95%E0%A5%89%E0%A4%B2%E0%A5%8B%E0%A4%A8%E0%A5%80_(%E0%A4%9C%E0%A5%80%E0%A4%B5_%E0%A4%AC%E0%A4%BF%E0%A4%9C%E0%A5%8D%E0%A4%9E%E0%A4%BE%E0%A4%A8)" title="कॉलोनी (जीव बिज्ञान) – Bhojpuri" lang="bh" hreflang="bh" class="interlanguage-link-target">भोजपुरी</A></LI><LI class="interlanguage-link interwiki-ca" style="display: none;"><A href="https://ca.wikipedia.org/wiki/Col%C3%B2nia_(biologia)" title="Colònia (biologia) – Catalan" lang="ca" hreflang="ca" class="interlanguage-link-target">Català</A></LI><LI class="interlanguage-link interwiki-cs" style="display: none;"><A href="https://cs.wikipedia.org/wiki/Kolonie_(biologie)" title="Kolonie (biologie) – Czech" lang="cs" hreflang="cs" class="interlanguage-link-target">Čeština</A></LI><LI class="interlanguage-link interwiki-de"><A href="https://de.wikipedia.org/wiki/Kolonie_(Biologie)" title="Kolonie (Biologie) – German" lang="de" hreflang="de" class="interlanguage-link-target">Deutsch</A></LI><LI class="interlanguage-link interwiki-et" style="display: none;"><A href="https://et.wikipedia.org/wiki/Koloonia_(bioloogia)" title="Koloonia (bioloogia) – Estonian" lang="et" hreflang="et" class="interlanguage-link-target">Eesti</A></LI><LI class="interlanguage-link interwiki-el" style="display: none;"><A href="https://el.wikipedia.org/wiki/%CE%91%CF%80%CE%BF%CE%B9%CE%BA%CE%AF%CE%B1_(%CE%BC%CE%B9%CE%BA%CF%81%CE%BF%CE%B2%CE%B9%CE%BF%CE%BB%CE%BF%CE%B3%CE%AF%CE%B1)" title="Αποικία (μικροβιολογία) – Greek" lang="el" hreflang="el" class="interlanguage-link-target">Ελληνικά</A></LI><LI class="interlanguage-link interwiki-es"><A href="https://es.wikipedia.org/wiki/Colonia_(biolog%C3%ADa)" title="Colonia (biología) – Spanish" lang="es" hreflang="es" class="interlanguage-link-target">Español</A></LI><LI class="interlanguage-link interwiki-eo" style="display: none;"><A href="https://eo.wikipedia.org/wiki/Kolonio_(biologio)" title="Kolonio (biologio) – Esperanto" lang="eo" hreflang="eo" class="interlanguage-link-target">Esperanto</A></LI><LI class="interlanguage-link interwiki-fr"><A href="https://fr.wikipedia.org/wiki/Colonie_(biologie)" title="Colonie (biologie) – French" lang="fr" hreflang="fr" class="interlanguage-link-target">Français</A></LI><LI class="interlanguage-link interwiki-ko"><A href="https://ko.wikipedia.org/wiki/%EA%B5%B0%EC%83%9D" title="군생 – Korean" lang="ko" hreflang="ko" class="interlanguage-link-target">한국어</A></LI><LI class="interlanguage-link interwiki-hy" style="display: none;"><A href="https://hy.wikipedia.org/wiki/%D4%B3%D5%A1%D5%B2%D5%B8%D6%82%D5%A9_(%D5%AF%D5%A5%D5%B6%D5%BD%D5%A1%D5%A2%D5%A1%D5%B6%D5%B8%D6%82%D5%A9%D5%B5%D5%B8%D6%82%D5%B6)" title="Գաղութ (կենսաբանություն) – Armenian" lang="hy" hreflang="hy" class="interlanguage-link-target">Հայերեն</A></LI><LI class="interlanguage-link interwiki-hr" style="display: none;"><A href="https://hr.wikipedia.org/wiki/Kolonija_(biologija)" title="Kolonija (biologija) – Croatian" lang="hr" hreflang="hr" class="interlanguage-link-target">Hrvatski</A></LI><LI class="interlanguage-link interwiki-is" style="display: none;"><A href="https://is.wikipedia.org/wiki/Bakter%C3%ADu%C3%BEyrping" title="Bakteríuþyrping – Icelandic" lang="is" hreflang="is" class="interlanguage-link-target">Íslenska</A></LI><LI class="interlanguage-link interwiki-it"><A href="https://it.wikipedia.org/wiki/Colonia_(zoologia)" title="Colonia (zoologia) – Italian" lang="it" hreflang="it" class="interlanguage-link-target">Italiano</A></LI><LI class="interlanguage-link interwiki-ku" style="display: none;"><A href="https://ku.wikipedia.org/wiki/Dagirker%C3%AE" title="Dagirkerî – Kurdish" lang="ku" hreflang="ku" class="interlanguage-link-target">Kurdî</A></LI><LI class="interlanguage-link interwiki-ky" style="display: none;"><A href="https://ky.wikipedia.org/wiki/%D0%9A%D0%BE%D0%BB%D0%BE%D0%BD%D0%B8%D1%8F%D0%BB%D1%83%D1%83_%D0%BE%D1%80%D0%B3%D0%B0%D0%BD%D0%B8%D0%B7%D0%BC%D0%B4%D0%B5%D1%80" title="Колониялуу организмдер – Kyrgyz" lang="ky" hreflang="ky" class="interlanguage-link-target">Кыргызча</A></LI><LI class="interlanguage-link interwiki-lt" style="display: none;"><A href="https://lt.wikipedia.org/wiki/Kolonija_(biologija)" title="Kolonija (biologija) – Lithuanian" lang="lt" hreflang="lt" class="interlanguage-link-target">Lietuvių</A></LI><LI class="interlanguage-link interwiki-nl" style="display: none;"><A href="https://nl.wikipedia.org/wiki/Kolonie_(biologie)" title="Kolonie (biologie) – Dutch" lang="nl" hreflang="nl" class="interlanguage-link-target">Nederlands</A></LI><LI class="interlanguage-link interwiki-no" style="display: none;"><A href="https://no.wikipedia.org/wiki/Koloni_(biologi)" title="Koloni (biologi) – Norwegian" lang="no" hreflang="no" class="interlanguage-link-target">Norsk</A></LI><LI class="interlanguage-link interwiki-pl" style="display: none;"><A href="https://pl.wikipedia.org/wiki/Kolonia_(biologia)" title="Kolonia (biologia) – Polish" lang="pl" hreflang="pl" class="interlanguage-link-target">Polski</A></LI><LI class="interlanguage-link interwiki-pt"><A href="https://pt.wikipedia.org/wiki/Col%C3%B3nia_(biologia)" title="Colónia (biologia) – Portuguese" lang="pt" hreflang="pt" class="interlanguage-link-target">Português</A></LI><LI class="interlanguage-link interwiki-ru"><A href="https://ru.wikipedia.org/wiki/%D0%9A%D0%BE%D0%BB%D0%BE%D0%BD%D0%B8%D1%8F_(%D0%B1%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F)" title="Колония (биология) – Russian" lang="ru" hreflang="ru" class="interlanguage-link-target">Русский</A></LI><LI class="interlanguage-link interwiki-simple" style="display: none;"><A href="https://simple.wikipedia.org/wiki/Colony_(biology)" title="Colony (biology) – Simple English" lang="simple" hreflang="simple" class="interlanguage-link-target">Simple English</A></LI><LI class="interlanguage-link interwiki-sh" style="display: none;"><A href="https://sh.wikipedia.org/wiki/Kolonija_(biologija)" title="Kolonija (biologija) – Serbo-Croatian" lang="sh" hreflang="sh" class="interlanguage-link-target">Srpskohrvatski / српскохрватски</A></LI><LI class="interlanguage-link interwiki-fi" style="display: none;"><A href="https://fi.wikipedia.org/wiki/Yhdyskunta_(biologia)" title="Yhdyskunta (biologia) – Finnish" lang="fi" hreflang="fi" class="interlanguage-link-target">Suomi</A></LI><LI class="interlanguage-link interwiki-sv" style="display: none;"><A href="https://sv.wikipedia.org/wiki/Koloni_(biologi)" title="Koloni (biologi) – Swedish" lang="sv" hreflang="sv" class="interlanguage-link-target">Svenska</A></LI><LI class="interlanguage-link interwiki-uk" style="display: none;"><A href="https://uk.wikipedia.org/wiki/%D0%9A%D0%BE%D0%BB%D0%BE%D0%BD%D1%96%D1%8F_(%D0%B1%D1%96%D0%BE%D0%BB%D0%BE%D0%B3%D1%96%D1%8F)" title="Колонія (біологія) – Ukrainian" lang="uk" hreflang="uk" class="interlanguage-link-target">Українська</A></LI><LI class="interlanguage-link interwiki-vi"><A href="https://vi.wikipedia.org/wiki/T%E1%BA%ADp_%C4%91o%C3%A0n_(sinh_h%E1%BB%8Dc)" title="Tập đoàn (sinh học) – Vietnamese" lang="vi" hreflang="vi" class="interlanguage-link-target">Tiếng Việt</A></LI>				<BUTTON class="mw-interlanguage-selector mw-ui-button" title="All languages (initial selection from common choices by you and others)">21 more</BUTTON></UL>
				<DIV class="after-portlet after-portlet-lang"><SPAN class="wb-langlinks-edit wb-langlinks-link"><A href="https://www.wikidata.org/wiki/Special:EntityPage/Q354341#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Edit links</A></SPAN></DIV>			</DIV>
		</DIV>
				</DIV>
		</DIV>
				<DIV id="footer" role="contentinfo">
						<UL id="footer-info">
								<LI id="footer-info-lastmod"> This page was last edited on 8 April 2018, at 21:15.</LI>
								<LI id="footer-info-copyright">Text is available under the <A rel="license" href="//en.wikipedia.org/wiki/Wikipedia:Text_of_Creative_Commons_Attribution-ShareAlike_3.0_Unported_License">Creative Commons Attribution-ShareAlike License</A><A rel="license" href="//creativecommons.org/licenses/by-sa/3.0/" style="display:none;"></A>;
additional terms may apply.  By using this site, you agree to the <A href="//wikimediafoundation.org/wiki/Terms_of_Use">Terms of Use</A> and <A href="//wikimediafoundation.org/wiki/Privacy_policy">Privacy Policy</A>. Wikipedia® is a registered trademark of the <A href="//www.wikimediafoundation.org/">Wikimedia Foundation, Inc.</A>, a non-profit organization.</LI>
							</UL>
						<UL id="footer-places">
								<LI id="footer-places-privacy"><A href="https://wikimediafoundation.org/wiki/Privacy_policy" class="extiw" title="wmf:Privacy policy">Privacy policy</A></LI>
								<LI id="footer-places-about"><A href="/wiki/Wikipedia:About" title="Wikipedia:About">About Wikipedia</A></LI>
								<LI id="footer-places-disclaimer"><A href="/wiki/Wikipedia:General_disclaimer" title="Wikipedia:General disclaimer">Disclaimers</A></LI>
								<LI id="footer-places-contact"><A href="//en.wikipedia.org/wiki/Wikipedia:Contact_us">Contact Wikipedia</A></LI>
								<LI id="footer-places-developers"><A href="https://www.mediawiki.org/wiki/Special:MyLanguage/How_to_contribute">Developers</A></LI>
								<LI id="footer-places-cookiestatement"><A href="https://wikimediafoundation.org/wiki/Cookie_statement">Cookie statement</A></LI>
								<LI id="footer-places-mobileview"><A href="//en.m.wikipedia.org/w/index.php?title=Colony_(biology)&mobileaction=toggle_view_mobile" class="noprint stopMobileRedirectToggle">Mobile view</A></LI>
							<LI style=""><A href="#">Enable previews</A></LI></UL>
										<UL id="footer-icons" class="noprint">
										<LI id="footer-copyrightico">
						<A href="https://wikimediafoundation.org/"><IMG src="/static/images/wikimedia-button.png" srcset="/static/images/wikimedia-button-1.5x.png 1.5x, /static/images/wikimedia-button-2x.png 2x" width="88" height="31" alt="Wikimedia Foundation" /></A>					</LI>
										<LI id="footer-poweredbyico">
						<A href="//www.mediawiki.org/"><IMG src="/static/images/poweredby_mediawiki_88x31.png" alt="Powered by MediaWiki" srcset="/static/images/poweredby_mediawiki_132x47.png 1.5x, /static/images/poweredby_mediawiki_176x62.png 2x" width="88" height="31" /></A>					</LI>
									</UL>
						<DIV style="clear: both;"></DIV>
		</DIV>

<DIV class="suggestions" style="display: none; font-size: 13px;"><DIV class="suggestions-results"></DIV><DIV class="suggestions-special"></DIV></DIV><DIV id="mwe-popups-svg"><svg xmlns="http://www.w3.org/2000/svg" width="0" height="0"><defs><clipPath id="mwe-popups-mask"><polygon points="0 8, 10 8, 18 0, 26 8, 1000 8, 1000 1000, 0 1000"></polygon></clipPath><clipPath id="mwe-popups-mask-flip"><polygon points="0 8, 274 8, 282 0, 290 8, 1000 8, 1000 1000, 0 1000"></polygon></clipPath><clipPath id="mwe-popups-landscape-mask"><polygon points="0 8, 174 8, 182 0, 190 8, 1000 8, 1000 1000, 0 1000"></polygon></clipPath><clipPath id="mwe-popups-landscape-mask-flip"><polygon points="0 0, 1000 0, 1000 242, 190 242, 182 250, 174 242, 0 242"></polygon></clipPath></defs></svg></DIV></BODY></HTML>

document_title: Colony (biology)

token: Colony

token: Biology

token: wikipedia

token: Jump

token: to

token: navigation

token: search

token: In

token: biology

token: a

token: colony

token: is

token: composed

token: of

token: two

token: or

token: more

token: conspecific

token: individuals

token: living

token: in

token: close

token: association

token: with

token: connected

token: one

token: another

token: This

token: usually

token: for

token: mutual

token: benefit

token: such

token: as

token: stronger

token: defense

token: the

token: ability

token: attack

token: bigger

token: prey

token: It

token: cluster

token: identical

token: cells

token: clones

token: on

token: surface

token: within

token: solid

token: medium

token: derived

token: from

token: single

token: parent

token: cell

token: bacterial

token: contrast

token: solitary

token: organism

token: which

token: all

token: live

token: independently

token: and

token: have

token: functions

token: needed

token: survive

token: српскохрватски

question_text: benefits of colonial life for single celled organisms